Anchusa officinalis

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Habitus and growth type

  • Height [m]: 0.2–1
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  • Growth form: monocarpic perennial non-clonal herb
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  • Life form: hemicryptophyte
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  • Life strategy: CS – competitor/stress-tolerator
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  • Life strategy (Pierce method based on leaf traits): CR
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  • Life strategy (Pierce method, C-score): 45.5 %
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  • Life strategy (Pierce method, S-score): 0 %
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  • Life strategy (Pierce method, R-score): 54.5 %
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Leaf

  • Leaf presence and metamorphosis: leaves present, not modified
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  • Leaf arrangement (phyllotaxis): alternate, rosulate
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  • Leaf shape: simple – entire
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  • Stipules: absent
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  • Petiole: both present and absent
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  • Leaf life span: summer green
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  • Leaf anatomy: scleromorphic, mesomorphic
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Flower

  • Flowering period [month]: May-September
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  • Flowering phase: 7 Ligustrum vulgare-Stachys sylvatica (end of early summer)
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  • Flower colour: white, blue-violet
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  • Flower symmetry: actinomorphic
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  • Perianth type: calyx and corolla
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  • Perianth fusion: fused
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  • Shape of the sympetalous corolla or syntepalous perianth: hypocrateriform
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  • Calyx fusion: synsepalous
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  • Inflorescence type: cincinnus
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  • Dicliny: synoecious, gynomonoecious, gynodioecious
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  • Generative reproduction type: mixed mating
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  • Pollination syndrome: insect-pollination, selfing
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Fruit, seed and dispersal

  • Fruit type: dry fruit – cluster of four one-seeded nutlets
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  • Fruit colour: brown
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  • Reproduction type: only by seed/spores
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  • Dispersal unit (diaspore): fruit, infrutescence or its part
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  • Dispersal strategy: Allium (mainly autochory)
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  • Myrmecochory: myrmecochorous
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Belowground organs and clonality

  • Shoot metamorphosis: pleiocorm
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  • Root metamorphosis: primary storage root
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  • Storage organ: pleiocorm, primary storage root
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  • Shoot life span (cyclicity): dicyclic or polycyclic shoots prevailing
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  • Primary root: present
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  • Position of root buds: primary root
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  • Role of root buds in life-history of a plant: regenerative
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  • Bud bank

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  • Number of buds per shoot at the soil surface (root buds excluded): 5
  • Number of buds per shoot at a depth of 0–10 cm (root buds excluded): 12
  • Number of buds per shoot at a depth greater than 10 cm (root buds excluded): 0
  • Size of the belowground bud bank (root buds excluded): 17
  • Depth of the belowground bud bank (root buds exluded) [cm]: 4
  • Number of buds per shoot at the soil surface (root buds included): 5
  • Number of buds per shoot at a depth of 0–10 cm (root buds included): 15
  • Number of buds per shoot at a depth greater than 10 cm (root buds included): 15
  • Size of the belowground bud bank (root buds included): 35
  • Depth of the belowground bud bank (root buds included) [cm]: 9

Trophic mode

  • Parasitism and mycoheterotrophy: autotrophic
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  • Carnivory: non-carnivorous
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  • Symbiotic nitrogen fixation: no nitrogen-fixing symbionts
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Karyology

  • Chromosome number (2n): 16
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  • Ploidy level (x): 2
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  • 2C genome size [Mbp]: 12908.71
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  • 1Cx monoploid genome size [Mbp]: 6454.35
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  • Genomic GC content: 38.5 %
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Taxon origin

  • Origin in the Czech Republic: archaeophyte
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  • Invasion status: naturalized
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  • Geographic origin: Europe, Mediterranean
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  • Introduction pathway: accidental introduction
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Ecological indicator values

  • Ellenberg-type indicator values

  • Light indicator value: 8 – light plant, only exceptionally occurring at less than 40% of diffuse radiation incident in an open area
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  • Temperature indicator value: 7 – heat indicator, occurring in relatively warm lowlands
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  • Moisture indicator value: 3 – missing on damp soil
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  • Reaction indicator value: 7 – indicator of slightly acidic to slightly basic conditions, never occurring in very acidic conditions
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  • Nutrient indicator value: 5 – occurring at moderately nutrient-rich sites, and less frequently at poor and rich sites
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  • Salinity indicator value: 0 – not salt tolerant, glycophyte
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Habitat and sociology

  • Occurrence in habitats

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  • 8 Dry grasslands

  • 8C Narrow-leaved sub-continental steppes: 1 – rare occurrence
  • 8E Acidophilous dry grasslands: 1 – rare occurrence
  • 9 Sand grasslands and rock-outcrop vegetation

  • 9C Festuca grasslands on acidic sands: 1 – rare occurrence
  • 13 Anthropogenic vegetation

  • 13D Perennial thermophilous ruderal vegetation: 2 – optimum
  • Affinity to the forest environment

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  • Affinity to the forest environment in Thermophyticum: 0 – taxon that does not spontaneously occur in Czech forests
  • Affinity to the forest environment in Mesophyticum and Oreophyticum: 0 – taxon that does not spontaneously occur in Czech forests
  • Ecological specialization indices

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  • Ecological specialization index for all vegetation types: 4
  • Ecological specialization index for non-forest vegetation: 3.8
  • Colonization ability

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  • Index of colonization success (ICS): 2
  • Index of colonization potential (ICP): 6
  • Optimum successional age [years]: 25

Distribution and frequency

  • Floristic zone: boreal, northern temperate, southern temperate, submeridional, meridional
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  • Floristic region:
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  • Continentality degree: 6
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  • Distribution range extension along the continentality gradient: 5
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  • Elevational belt in the Czech Republic: lowlands, colline belt, submontane belt
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  • Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells: ,
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  • Commonness in vegetation plots from the Czech Republic

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  • Occurrence frequency in vegetation plots: 0.1 %
  • Occurrence frequency in vegetation plots with a cover above 5%: 11.1 %
  • Occurrence frequency in vegetation plots with a cover above 25%: 0 %
  • Occurrence frequency in vegetation plots with a cover above 50%: 0 %
  • Mean percentage cover in vegetation plots: 2.7 %
  • Maximum percentage cover in vegetation plots: 13 %
  • Number of habitats with taxon occurrence in the Czech Republic

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  • Number of narrow habitats in which the taxon occurs: 4
  • Number of narrow habitats in which the taxon has its optimum: 1
  • Number of broad habitats in which the taxon occurs: 3
  • Number of broad habitats in which the taxon has its optimum: 1

Threats and protection

  • Red List 2017 (national categories): taxon is not on the Red List
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  • Red List 2017 (IUCN categories): LC(NA) – least concern (taxon is not on the Red List)
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  • Legal protection: not protected by law
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