Betula pendula var. obscura

Leaf

Leaf presence and metamorphosis: leaves present, not modified

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Leaf presence and metamorphosis
Data on the presence of leaves on the plant, their metamorphoses and reductions are based on the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010) and the Key to the Flora of the Czech Republic (Kubát et al. 2002).

Categories

leaves present, not modified

leaves modified to spines

leaves modified to tendrils

leaves modified to phyllodes

leaves modified to pitchers

leaves reduced to collars

leaves reduced to sheaths

leaves reduced to scales

leaves absent

Data source and citation

Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Kubát K., Hrouda L., Chrtek J. Jr., Kaplan Z., Kirschner J. & Štěpánek J. (eds) (2002) Klíč ke květeně České republiky [Key to the flora of the Czech Republic]. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Leaf arrangement (phyllotaxis): alternate

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Leaf arrangement (phyllotaxis)
Four basic types of leaf arrangement are distinguished: alternate, opposite, verticillate (whorled) and rosulate (in the basal rosette). The character is assessed in well-developed plants, i.e. not in individuals re-sprouting after damage by mowing or grazing or those with teratological modifications. More than one character state may occur (e.g. Hylotelephium jullianum and Salix purpurea) in some taxa: all character states are recorded in such cases.

In some plants, the arrangement of frondose bracts in the inflorescence is assessed separately (e.g. true leaves in Veronica persica and V. polita are opposite, while bracts are alternate). Leaves with interpetiolar stipules found in the Rubiaceae family are considered as whorled. The leaves in Rhamnus cathartica are considered as opposite, although in most cases they are sub-opposite.

The information was extracted mainly from the descriptions in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). In cases of uncertainties, mainly for alien taxa, additional sources were consulted, including the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994) and the Flora of Pakistan (www.tropicos.org/Project/Pakistan).

Categories

alternate

opposite

verticillate

rosulate

Data source and citation

Grulich V., Holubová D., Štěpánková P. & Řezníčková M. (2017) Leaf arrangement. – www.pladias.cz.

Further references

Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.
Leaf shape: simple – entire

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Leaf shape
The primary distinction is made between simple and compound leaves. The simple leaves are categorized based on the leaf blade division associated with venation into palmately divided (e.g. Alchemilla), pinnately divided (e.g. Achillea millefolium), forked (e.g. Batrachium, Ceratophyllum and Utricularia) and pedate (e.g. Helleborus). The categorization is based on well-developed leaves. In many taxa, transitions occur between simple leaves with a dentate or serrate margin, and simple divided (pinnately or palmately lobed) leaves. Only the leaves with the lamina divided to at least one-quarter of their width are considered as divided. Many taxa with varying leaf division are assigned to more than one character state.

The compound leaves are divided into palmate and pinnate. The taxa that have both ternate and pinnate leaves, the latter with two pairs of leaflets (e.g. Aegopodium podagraria and some other species of the Apiaceae family), are assigned to both character states. The degree of division in pinnately compound leaves indicated here relates to well-developed leaves, especially to the basal part of the lamina. Taxa with multiple pinnately compound leaves are assigned to two or more character states based on the level of division, but very small leaves, which may correspond to simple leaves, are not considered.

In many cases, there are transitions between simple and compound leaves, especially between pinnatisect and pinnate leaves. Leaves with linear or filiform segments, including the bi-, tri- or even more-pinnatisect or palmatisect leaves (e.g. stem leaves in Batrachium fluitans, Cardamine pratensis and the genus Seseli) are classified as simple (dissected) leaves. In contrast, leaves with broader segments attached to the rachis by a distinct constriction or a petiolule (e.g. stem leaves in Cardamine dentata or ground leaves in Pimpinella saxifraga) are classified as compound.

In heterophyllous taxa, all types of leaves are assessed, and the taxon is assigned to two or more character states. However, less divided leaves found in juvenile plants of some taxa are not considered heterophyllous. The parasitic plants with rudimentary (vestigial) leaves (e.g. Cuscuta) or the plants with phylloclades replacing the vestigial leaves (e.g. Asparagus) are assigned the character state “reduced”.

The information was extracted mainly from the descriptions in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). In uncertain cases, mainly for alien taxa, additional sources were consulted, including the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994) and the Flora of Pakistan (www.tropicos.org/Project/Pakistan).

Categories

simple – entire

simple – palmately divided

simple – pinnately divided

simple – forked

simple – pedate

compound – ternate

compound – palmate (5-foliate)

compound – palmate (7-foliate)

compound – palmate (8- and more-foliate)

compound – imparipinnate

compound – paripinnate

compound – interruptedly pinnate

compound – bipinnate

compound – tripinnate

compound – quadripinate

reduced

Data source and citation

Grulich V., Holubová D., Štěpánková P. & Řezníčková M. (2017) Leaf shape. – www.pladias.cz.

Further references

Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.
Stipules: present

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Stipules
Stipules, i.e. paired leaflike appendages at the base of the petiole or sessile leaf blade, can be present or absent. Caducous stipules, i.e. those disappearing soon after the leaf blade has developed (e.g. Prunus), are considered as present. The interpetiolar stipules, morphologically indistinguishable from true leaves and together forming whorls (e.g. Rubiaceae), are considered as true stipules. In contrast, stipules modified into glands (e.g. Lotus) or hairs (e.g. Portulacaceae) are not considered as stipules here.

Information about the presence of stipules was extracted from the descriptions in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). In cases of uncertainties, mainly concerning alien taxa, descriptions in the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994) and the Flora of Pakistan (www.tropicos.org/Project/Pakistan) were consulted.

Categories

present

absent

Data source and citation

Grulich V., Holubová D., Štěpánková P. & Řezníčková M. (2017) Stipules. – www.pladias.cz.

Further references

Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.
Petiole: present

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Petiole
Leaf petiole can be present or absent. In some plants, it can be present in some leaves but absent in others. The data were extracted from the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010), the Key to the Flora of the Czech Republic (Kubát et al. 2002), the New Hungarian Herbal (Király et al. 2011) and the Excursion Flora of Germany (Jäger & Werner 2000).

Categories

present

mainly present

both present and absent

mainly absent

absent

Data source and citation

Prokešová H. & Grulich V. (2017) Petiole. – www.pladias.cz.

Further references

Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Jäger E. J. & Werner K. (eds) (2000) Rothmaler, Exkursionsflora von Deutschland. Band 3. Gefäßpflanzen: Atlasband. Ed. 10. – Spectrum Akademischer Verlag, Heidelberg & Berlin.
Király G., Virók V. & Molnár V. (eds) (2011) Új Magyar füvészkönyv. Magyarország hajtásos növényei: ábrák [New Hungarian Herbal. The vascular plants of Hungary: Figures]. – Aggteleki Nemzeti Park Igazgatóság, Jósvafő.
Kubát K., Hrouda L., Chrtek J. Jr., Kaplan Z., Kirschner J. & Štěpánek J. (eds) (2002) Klíč ke květeně České republiky [Key to the flora of the Czech Republic]. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.

Flower

Flower colour: yellow-green

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Flower colour
Flower colour is reported for nearly all angiosperms except duckweeds (Araceae p. p.) and some hybrids for which data on flower colour were not available.

If a species has more than one flower colour, all colours are reported irrespective of their frequency. This approach is used both for species that regularly form populations with different flower colours (e.g. Corydalis cava and Iris pumila) and for species with occasional occurrence of deviating flower colour (e.g. albinism in Salvia pratensis or pink flowers in Ajuga reptans). However, the whole range of variation is not fully reported in cultivated plants, for which some cultivars of different colour may be ignored (e.g. Gladiolus hortulanus and Callistephus chinensis). In plants with flowers of two colours (e.g. Cypripedium calceolus), both colours are reported. In plants with multi-coloured flowers (e.g. the variegated lip in Ophrys apifera) the predominant colour is reported.

If the flower has a well-developed perianth, the reported flower colour relates to the corolla or the tepals of the homochlamydeous perianth. If such a flower has bracts of a contrasting colour (e.g. Melampyrum nemorosum), their colour is not considered. If the corolla or the homochlamydeous perianth is not developed, the flower colour is based on the calyx (e.g. Daphne mezereum), bracts (e.g. Aristolochia clematitis), the system of bracts and bracteoles in the inflorescence (Euphorbia) or the involucre on secondary peduncles (Bupleurum longifolium). In species of Araceae with spadix and spathe of contrasting colours (e.g. Calla palustris) both colours are reported. The colour of the whole inflorescence is reported for some plants with reduced flowers (e.g. Betula, Salix, some Cyperaceae and Typhaceae). Spikelets in Poaceae are reported as green disregarding a possible violet tint; exceptions include the Melica ciliata agg. and Cortaderia that are reported as white. Also in other, rare cases, the inflorescence colour is reported as flower colour (e.g. green in Ficus carica). In Asteraceae, the colours of the disk flowers and ray flowers are reported separately if the ray flowers are developed and have a contrasting colour (e.g. Bellis perennis). The colour of the involucrum is reported for species with tiny flower heads and indistinct flowers (e.g. Artemisia campestris and Xanthium) and for “immortelles” (e.g. Helichrysum and Xeranthemum).

Information on flower colour is partly based on the field knowledge, partly obtained from various photographs and descriptions in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). In the taxa that are not reported in the Flora of the Czech Republic, as well as in unclear cases (especially in alien species), other sources were used, especially the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994) and the Flora of Pakistan (http://www.tropicos.org/Project/Pakistan).

Categories

white (including grey and silvery, and rare cases of individuals with white colour) – e.g. ray flowers of Leucanthemum vulgare, albinotic plants of Glechoma hederacea, catkins of Salix caprea

yellow-white (including white-yellow and cream) – e.g. Scabiosa ochroleuca

green-white (including white-green and greenish) – e.g. Orthilia secunda

green – e.g. Poa pratensis (the colour relates to the glumes and lemmas)

yellow-green (including green-yellow) – e.g. Acer pseudoplatanus, Rhamnus cathartica

yellow – e.g. Taraxacum

orange – e.g. Pilosella aurantiaca

pink (including white-pink, pink-white and dark pink) – e.g. Malva alcea, Rosa canina

pink-violet – e.g. Allium schoenoprasum

red – e.g. Papaver rhoeas

red-violet (includes all the hues of purple, pink-red and violet-red) – e.g. Astragalus onobrychis, Cirsium palustre

violet (including dark violet and black-violet) – e.g. Bartsia alpina, Salvia verticillata

blue – e.g. Centaurea cyanus

blue-violet – e.g. Aconitum plicatum

brown (including yellow-brown, brown-yellow, beige and brown-violet) – e.g. Euonymus verrucosus, Neottia nidus-avis

red-brown – e.g. Asarum europaeum, Scrophularia nodosa

black – e.g. Carex acuta (the colour relates to the colour of bracts)

Categories

white

yellow-white

green-white

green

yellow-green

yellow

orange

pink

pink-violet

red

red-violet

violet

blue

blue-violet

brown

red-brown

black

Data source and citation

Štěpánková P. & Grulich V. (2019) Flower colour. – www.pladias.cz.

Further references

Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.
Perianth type: homochlamydeous, reduced or absent

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Perianth type
Perianth (perigon), i.e. the non-reproductive part of the angiosperm flower, can be classified into heterochlamydeous and homochlamydeous. Heterochlamydeous flowers are divided into calyx and corolla. In homochlamydeous flowers, calyx and corolla are indistinguishable. Perianth or some of its parts can be reduced or absent; flowers with no perianth are called achlamydeous.

In Apiaceae, the presence of the calyx teeth is assessed as a reduced calyx; if these teeth are not visible, the calyx is considered as absent. In Asteraceae, the presence of a pappus, scales or a collar-like structure is considered as a reduced calyx; if no such structures are present, the calyx is considered as absent. In Cyperaceae, the presence of perianth bristles is assessed as a reduced perianth. All members of the Poaceae family are considered as plants with a reduced perianth. The perianth in the genus Basella is arbitrarily classified as a reduced calyx though it is also often considered as a reduced homochlamydeous perianth. The character states “homochlamydeous, sometimes absent” and “homochlamydeous, reduced or absent” mean that in one plant some flowers may have a well-developed or reduced perianth, while other flowers may be achlamydeous (e.g. Atriplex).

The information was extracted mainly from the descriptions in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). For the taxa not treated in that flora or if uncertainties occurred, mainly concerning some alien taxa, the descriptions in the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994) and the Flora of Pakistan (www.tropicos.org/Project/Pakistan) were consulted.

Categories

homochlamydeous

homochlamydeous, sometimes reduced

homochlamydeous, sometimes reduced or absent

homochlamydeous, sometimes absent

homochlamydeous, reduced or absent

calyx and corolla

calyx and corolla, corolla reduced or absent

calyx and corolla, corolla sometimes absent

calyx present, corolla sometimes reduced

calyx present, corolla reduced

calyx present, corolla reduced or absent

calyx present, corolla sometimes absent

calyx present, corolla absent

calyx reduced, corolla present

calyx sometimes absent, corolla present

calyx absent, corolla present

calyx absent, corolla sometimes present

reduced

reduced or absent

flower achlamydeous

Data source and citation

Grulich V., Prokešová H. & Štěpánková P. (2017) Perianth type. – www.pladias.cz.

Further references

Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.
Perianth fusion: reduced

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Perianth fusion
This characteristic of angiosperm flowers is assessed either as a fusion of the corolla or, in homochlamydeous taxa (e.g. Amaryllidaceae, Liliaceae and Orchidaceae) as a fusion of the whole perianth. It is not assessed in achlamydeous groups (e.g. Salix) and plants with a strongly reduced or rudimentary perianth or with the perianth modified in scale-like or setaceous structures with a varying number of bristles, which may be free (e.g. in Cyperaceae) or partially fused (e.g. in most of Poaceae). The perianth of such plants is considered as reduced. The perianth in the genus Aristolochia is also classified as reduced (neither fused nor free): it is modified to scales situated at the bottom of a tube-like structure formed by fused bracts. Both primary character states are assigned to the taxa with unisexual male and female flowers that differ in the fusion of the perianth (e.g. Cannabis). A similar approach is used in the taxa in which some flowers are homochlamydeous while others are achlamydeous (e.g. Atriplex).

The basic information was extracted from the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). If some uncertainty occurred, especially for alien taxa, other sources were consulted, including the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994) and the Flora of Pakistan (www.tropicos.org/Project/Pakistan).

Categories

free

fused

reduced

Data source and citation

Grulich V., Holubová D., Štěpánková P. & Řezníčková M. (2017) Perianth fusion. – www.pladias.cz.

Further references

Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.
Inflorescence type: amentum e floribus masculis, amentum e floribus femineis

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Inflorescence type
Inflorescence types follow the morphological system used in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). As the Czech terminology used for inflorescences does not match the English terminology, we use Latin terms in the English version of the Pladias Database. The exact identification of the inflorescence type is often equivocal because of varying interpretations of the same object. In species with unisexual flowers, male and female flowers can occur in different inflorescence types. In other cases, it is not possible to identify the inflorescence without detailed knowledge of evolutionary morphology, e.g. umbella vs pseudumbella in the genus Butomus. There are also compound inflorescences, in some cases with very different structure of their parts, especially in Asteraceae, which can have even triple inflorescences (e.g. Echinops sphaerocephalus often has an anthella ex capitulis anthodiorum composita).

The information was extracted mainly from the descriptions in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). For the taxa not treated in that flora or if some uncertainties occurred, mainly concerning some alien taxa, information was taken from the descriptions in the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994) and the Flora of Pakistan (www.tropicos.org/Project/Pakistan). In critical groups (e.g. Rubus), especially in recently described species, inflorescence type was taken from the original descriptions.

Categories

pseudumbella e cyathiis composita

capitulum

capitulum e floribus masculis compositum

capitulum e spiculis compositum

capitulum e verticillastris compositum

capitulum e floribus femineis compositum

capitulum ex anthodiis compositum

racemus

racemus e capitulis compositus

racemus ex corymbis anthodiorum compositus

racemus e spiculis compositus

racemus e verticillastris compositus

racemus ex umbellis compositus

racemus e floribus masculis compositus

racemus e floribus femineis compositus

racemus e fasciculis compositus

racemus ex anthodiis compositus

racemus ex anthodiis masculis compositus

racemus e cincinnis compositus

corymbothyrsus

corymbothyrsus e fasciculis compositus

corymbothyrsus ex fasciculis anthodiorum compositus

corymbothyrsus ex anthodiis compositus

corymbus

corymbus ex anthodiis compositus

amentum

amentum e floribus masculis

amentum e floribus femineis

spica

spica e spiculis composita

spica e floribus masculis composita

spica e floribus femineis composita

spicula

anthella

anthella ex capitulis anthodiorum composita

anthella e spiculis composita

anthella e floribus masculis composita

anthella ex anthodiis composita

flores solitarii

flores solitarii masculi

flores solitarii feminei

panicula

panicula e capitulis composita

panicula ex capitulis anthodiorum composita

panicula e corymbis composita

panicula e spiculis composita

panicula e spiculis masculis composita

panicula e pseudospicis composita

panicula e floribus masculis composita

panicula e fasciculis composita

panicula e bostrychibus composita

panicula ex anthodiis composita

panicula e dichasiis composita

panicula e cincinnis composita

pseudospica

pseudospica e capitulis composita

pseudospica e spiculis composita

pseudospica e verticillastris composita

pseudospica e floribus masculis composita

pseudospica ex anthodiis composita

pseudumbella

pseudumbella ex anthodiis composita

verticillastrum

pseudoracemus

umbella

umbella e spicis spicularum composita

spadix

spadix e floribus femineis compositus

umbella composita

fasciculus

fasciculus e floribus masculis compositus

fasciculus e floribus femineis compositus

fasciculus ex anthodiis femineis compositus

syconium

strobilus

bostryx

anthodium

anthodium solitarium

rhipidium

dichasium

dichasium e floribus femineis compositum

dichasium ex anthodiis compositum

cincinnus

Data source and citation

Grulich V. & Štěpánková P. (2019) Inflorescence type. – www.pladias.cz.

Further references

Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.

Fruit, seed and dispersal

Fruit type: dry fruit – achene/cypsela/samara

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Fruit type
The primary classification of fruit types is into dry and fleshy. Within each of these two groups, fruit types are further classified based on the scheme outlined in the first volume of the Flora of the Czech Republic (Slavíková 1988), which consistently uses the typological method. This means that fruits are classified based purely on their morphology following the formal definitions of the fruit type, regardless of the fruit type found in closely related species or genera.

One-seeded fruits in Brassicaceae (e.g. Crambe) are classified as achenes, not siliculas. Indehiscent two- and more-seeded fruits in the same family, breaking mainly in constrictions (e.g. in Bunias and Raphanus), are consistently classified as a loment, even if the fruit breaks into two distinct parts, of which one is one-seeded and the other, of strikingly different shape, two- or more-seeded and dehiscent, such as in Rapistrum rugosum. A similar approach is used for the classification of fruits in Fabaceae. Dehiscent fruits of most taxa are classified as legumes, while indehiscent two- and more-seeded fruits breaking into single-seeded parts (e.g. in Hippocrepis and Securigera) are classified as loments. One-seeded indehiscent fruits (e.g. in Onobrychis and Trifolium) are classified as achenes. Two- or more-seeded indehiscent fruits (e.g. in Sophora japonica and Vicia faba) are also classified as legumes. The fruits of all Euphorbia species are classified as capsules, although in some cases the seeds are not released. Fleshy false fruits of the genera Basella, Ficus, Maclura, Morus, Nuphar and Nymphaea are merged into a separate category.

The information about fruit type was extracted mainly from the descriptions in the Flora of the Czech Republic (vols. 1–8; Hejný et al. 1988–1992, Slavík et al. 1997–2004, Štěpánková et al. 2010). For the taxa not treated in that flora or in case of uncertainties, especially regarding alien taxa, descriptions in the Flora of North America (Flora of North America Editorial Committee 1993), the Flora of China (Wu et al. 1994), the Flora of Pakistan (www.tropicos.org/Project/Pakistan), and Flora Iberica (Castroviejo et al. 1986; mainly for the Fabaceae family) were consulted.

Categories

dry fruit – disciform breaking into mericarps

dry fruit – pair of nutlets

dry fruit – cremocarp

dry fruit – double samara

dry fruit – legume

dry fruit – follicle

dry fruit – achene/cypsela/samara

dry fruit – nut enclosed in an utricle

dry fruit – caryopsis

dry fruit – nut

dry fruit – schizocarp

dry fruit – cluster of follicles

dry fruit – head of achenes

dry fruit – lomentum/transversely dehiscent siliqua

dry fruit – siliqua

dry fruit – silicula

dry fruit – capsule

dry fruit – cluster of four one-seeded nutlets

dry fruit – dry schizocarp with an apical beak

fleshy fruit – berry

fleshy fruit – pome

fleshy fruit – drupe

fleshy fruit – head of one-seeded drupes

fleshy fruit – hip

fleshy fruit – special type

Data source and citation

Grulich V., Holubová D., Štěpánková P. & Řezníčková M. (2017) Fruit type. – www.pladias.cz.

Further references

Castroviejo S., Laínz M., López González G., Montserrat P., Muńoz Garmendia F., Paiva J. & Villar L. (eds) (1986) Flora Iberica. Plantas vasculares de la Península Ibérica e Islas Baleares. – Real Jardín Botánico, Madrid.
Flora of North America Editorial Committee (eds) (1993) Flora of North America North of Mexico. – Oxford University Press, New York.
Flora of Pakistan. – http://www.tropicos.org/Project/Pakistan
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Slavíková Z. (1988) Terminologický slovník [Terminological dictionary]. – In: Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds), Květena České socialistické republiky [Flora of the Czech Socialist Republic] 1: 130–153, Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Wu Z., Raven P. H. & Huang D. (eds) (1994) Flora of China. – Science Press, Beijing & Missouri Botanical Garden, St. Louis.
Dispersal unit (diaspore): seed

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Dispersal unit (diaspore)
Diaspore, also called dispersule or propagule, is a generative or vegetative part of the plant body that is dispersed from the parental plant and can produce a new individual. Generative diaspores include spores, seeds and fruits or similar dispersal units (e.g. aggregate fruits in Fragaria, multiple fruits in Morus, gymnosperm cones, epimatium-bearing seed in Taxus, spikelets or their various fragments in Poaceae). If the seed is released from dehiscent fruit or decaying ripe fleshy fruit, both seed and fruit can be considered as diaspores. In plants with indehiscent fruits, only the fruit is considered as a diaspore. A specific category of generative diaspore is tumbleweeds, i.e. mature plant parts including stem branches and large inflorescence (e.g. Crambe tataria and Falcaria vulgaris).

Vegetative diaspores are viable and movable parts of plants that originate above ground or in water and disconnect from the parent plant before sprouting. We did not consider as vegetative diaspores clonal organs connected with the maternal plant until the new plant becomes independent (e.g. stolons in Fragaria) and various types of below-ground organs or shoot bases embedded in soil (e.g. tubers of Helianthus tuberosus or grass tillers). Vegetative diaspores include (i) turions (e.g. Myriophyllum and Utricularia) and similar overwintering structures (detachable buds in Elodea and Groenlandia and shortened shoots of some pondweeds produced by rhizome or stolon, e.g. Potamogeton alpinus); (ii) bulbils and tubers of stem origin (e.g. Allium oleraceum and Dentaria bulbifera) or root origin (Ficaria only); (iii) plantlets born by pseudovivipary (e.g. Poa alpina); (iv) plantlets born from buds on leaves (e.g. Cardamine pratensis); (v) plantlets born on free ends of stolons, detachable before establishing (e.g. Hydrocharis and Jovibarba); (vi) unspecialized fragments of the shoot (e.g. Sedum album and many aquatic plants), shoot tips (e.g. Ceratophyllum demersum) or detachable offsprings born from axillary buds (e.g. Agrostis canina, Arabidopsis halleri and Rorippa amphibia); (vii) budding plants (Lemnaceae only); and (viii) gemmae produced by gametophytes (Trichomanes speciosum only).

Categories

spore

seed

fruit, infrutescence or its part

tumbleweed

turion

bulbil or tuber

pseudovivipary

leaf-born plantlet

stolon-born plantlet

shoot fragment

budding

gametophyte-gemma

Data source and citation

Sádlo J., Chytrý M., Pergl J. & Pyšek P. (2018) Plant dispersal strategies: a new classification based on themultiple dispersal modes of individual species. – Preslia 90: 1–22.
Dispersal strategy: Epilobium (mainly anemochory and autochory)

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Dispersal strategy
Plants use different dispersal modes, also called dispersal syndromes, depending on different dispersal vectors. For example, anemochory is the dispersal by wind, hydrochory by water, epizoochory by attachment to an animal body and endozoochory by animals via ingestion. However, single plant species usually use a combination of several dispersal modes rather than a single mode. Distinct combinations of dispersal modes repeatedly occurring in different plant taxa are called dispersal strategies. Sádlo et al. (2018) distinguished nine dispersal strategies named for the genus names of typical representatives. Taxa of the Czech flora are assigned to individual strategies based on this source.

Categories

Allium type – mainly autochory, less frequently anemochory, endozoochory and epizoochory. This is the most common dispersal strategy, including about 56% taxa of the Czech flora. About half of the included taxa are dispersal generalists lacking a clear morphological indication of anemochory or zoochory. Most myrmecochorous or probably myrmecochorous species are also assigned to this category.

Bidens type – mainly autochory and epizoochory, less frequently endozoochory. This dispersal strategy is characterized by two essential dispersal modes, of which autochory is the more important, despite the presence of morphological structures indicating epizoochory.

Cornus type – mainly autochory and endozoochory. Herbaceous plants, shrubs and small trees with fleshy fruit, often of the Rosaceae family, typically have this strategy. Furthermore, tall trees bearing large, heavy and nutrient-rich seeds are also included.

Epilobium type – mainly anemochory and autochory, less frequently endozoochory and epizoochory. This dispersal strategy is typical of taxa growing in mesic and dry habitats.

Lycopodium type – mainly anemochory, less frequently autochory, endozoochory, epizoochory and hydrochory. This dispersal strategy relies on light, very small spores and seeds that are dispersed, besides wind, by a wide range of vectors. Compared to other strategies, the role of autochory is small.

Phragmites type – mainly anemochory and hydrochory, less frequently autochory, endozoochory and epizoochory. Wetland taxa with light diaspores (both seeds and fruits) equipped with a hairy flying apparatus. Most of the taxa with this dispersal strategy lack vegetative diaspores. Woody plants, stout clonal graminoids and herbaceous plants are typical growth forms associated with this dispersal strategy.

Sparganium type – mainly autochory and hydrochory, less frequently endozoochory and epizoochory. This dispersal strategy is a wetland analogue of the Wolffia type, assigned to aquatic plants. It applies mainly to monocotyledonous taxa producing achenes with good buoyancy and with vegetative diaspores having an important role.

Wolffia type – mainly hydrochory, less frequently endozoochory and epizoochory. This dispersal strategy is typical of aquatic macrophytes spread by fruit, seed or spores. However, vegetative reproduction dominates in most cases, including stem fragmentation, the formation of stolons or, in Lemnaceae, budding colonies.

Zea type. Taxa with this dispersal strategy rarely or never disperse by generative diaspores and do not form vegetative aboveground diaspores.

Categories

Allium (mainly autochory)

Bidens (mainly autochory and epizoochory)

Cornus (mainly autochory and endozoochory)

Epilobium (mainly anemochory and autochory)

Lycopodium (mainly anemochory)

Phragmites (mainly anemochory and hydrochory)

Sparganium (mainly autochory and hydrochory)

Wolffia (mainly hydrochory)

Zea (no dispersal)

Data source and citation

Sádlo J., Chytrý M., Pergl J. & Pyšek P. (2018) Plant dispersal strategies: a new classification based on themultiple dispersal modes of individual species. – Preslia 90: 1–22.
Myrmecochory: non-myrmecochorous (b)

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Myrmecochory
Myrmecochorous plants, i.e. taxa dispersed by ants, possess an elaiosome, a nutrient-rich fleshy appendage of seed or fruit. However, in many taxa, the morphological indication or direct evidence of myrmecochory is equivocal. Removal experiments (seeds with and without elaiosome offered to ants) or chemical analysis (different nutrient content between seed and elaiosome; Konečná et al. 2018) would be needed to decide whether the appendage is elaiosome or not. Therefore, more categories than a simple binary distinction between myrmecochorous and non-myrmecochorous are recognized here:

myrmecochorous – an elaiosome (a conspicuous fleshy appendage of seed or fruit) is present

probably myrmecochorous – a fleshy structure resembling an elaiosome is present, but the appendage:seed size ratio is very small so that the possible advantage of seed transport for the ants would be small

probably non-myrmecochorous – seed or fruit bears a structure not similar to elaiosome, e.g. a non-fleshy appendage or rim

non-myrmecochorous (a) – an elaiosome is not present although the taxon is reported as myrmecochorous in the literature or is a close relative of taxa considered as myrmecochorous

non-myrmecochorous (b) – elaiosome is not present, and the taxon does not belong to a family reported as containing myrmecochorous species in the literature

Plant taxa that are often carried by ants to the nest although having no elaiosome (e.g. cheaters in this plant-ant mutualism or plant parts used as a building material for ant hills) are classified as non-myrmecochorous.

The data are based on the literature search and examination of seed samples of the taxa that are reported as myrmecochorous and their closely related congenerics. The list of these taxa with seed images is available at http://botanika.prf.jcu.cz/myrmekochorie/. These taxa were selected from the families represented in the Czech flora that contain at least one taxon reported as myrmecochorous in the literature (Sernander 1906, Hejný et al. 1988 onwards, Fitter & Peat 1994, Klotz et al. 2002, Grime et al. 2007, Kleyer et al. 2008, Servigne 2008, Lengyel et al. 2010, Študent 2012). Such taxa were found in 37 families including Amaryllidaceae, Apiaceae, Apocynaceae, Aristolochiaceae, Asparagaceae, Asteraceae, Boraginaceae, Campanulaceae, Caryophyllaceae, Celastraceae, Colchicaceae, Crassulaceae, Cyperaceae, Dipsacaceae, Euphorbiaceae, Fabaceae, Iridaceae, Juncaceae, Lamiaceae, Liliaceae, Linaceae, Montiaceae, Orobanchaceae, Oxalidaceae, Papaveraceae, Plantaginaceae, Poaceae, Polygalaceae, Polygonaceae, Portulacaceae, Primulaceae, Ranunculaceae, Resedaceae, Rosaceae, Santalaceae, Urticaceae and Violaceae. All the taxa not belonging to these families were classified as non-myrmecochorous (b).

For each of the five categories, a subcategory nv (= non vidimus, i.e. not seen) is used in the taxa for which we found neither information in the literature nor a photograph of a seed, and failed to collect seeds from living plants, but the assignment to the category is likely based on the traits of closely related taxa. For example, we have no data for Centaurea bruguiereana but we classify it as myrmecochorous nv, because all the taxa of Centaurea for which we have data possess an elaiosome.

Categories

myrmecochorous

myrmecochorous nv

probably myrmecochorous

probably myrmecochorous nv

probably non-myrmecochorous

probably non-myrmecochorous nv

non-myrmecochorous (a)

non-myrmecochorous (a) nv

non-myrmecochorous (b)

non-myrmecochorous (b) nv

Data source and citation

Konečná M., Štech M. & Lepš J. (2018) Myrmecochory. – www.pladias.cz.

Further references

Fitter A. H. & Peat H. J. (1994) The Ecological Flora Database. – Journal of Ecology 82: 415–425.
Grime J. P., Hodgson J. G. & Hunt R. (eds) (2007) Comparative plant ecology: a functional approach to common British species. 2nd edition. – Castlepoint Press, Colvend, Dalbeattie.
Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Kleyer M., Bekker R. M., Knevel I. C., Bakker J. P., Thompson K., Sonnenschein M., Poschlod P., van Groenendael J. M., Klimeš L., Klimešová J., Klotz S., Rusch G. M., Hermy M., Adriaens D., Boedeltje G., Bossuyt B., Dannemann A., Endels P., Götzenberger L., Hodgson J. G., Jackel A. K., Kühn I., Kunzmann D., Ozinga W. A., Romermann C., Stadler M., Schlegelmilch J., Steendam H. J., Tackenberg O., Wilmann B., Cornelissen J. H. C., Eriksson O., Garnier E. & Peco B. (2008) The LEDA Traitbase: a database of life-history traits of the Northwest European flora. – Journal of Ecology 96: 1266–1274.
Klotz S., Kühn I. & Durka W. (eds) (2002) BIOLFLOR: eine Datenbank mit biologisch-ökologischen Merkmalen zur Flora von Deutschland. – Schriftenreihe für Vegetationskunde 38: 1–334.
Konečná M., Moos M., Zahradníčková H., Šimek P. & Lepš J. (2018) Tasty rewards for ants: differences in elaiosome and seed metabolite profiles are consistent across species and reflect taxonomic relatedness. – Oecologia 188: 753–764.
Sernander R. (1906) Entwurf einer Monographie der europäischen Myrmekochoren. – Kungliga Svenska Vetenskapsakademiens Handlingar 41: 1–410.
Servigne P. (2008) Etude expérimentale et comparative de la myrmécochorie: le cas des fourmis dispersatrices Lasius niger et Myrmica rubra. – PhD thesis, Université libre de Bruxelles, Bruxelles.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Študent V. (2012) Společné funkční vlastnosti myrmekochorních druhů rostlin České republiky a sezónní a denní dynamika odnosu diaspor všivce lesního (Pedicularis sylvatica) mravenci [Traits of myrmecochorous plants of the Czech Republic and a seasonal and daily seed’s removal dynamics of lousewort (Pedicularis sylvatica) by ants]. – Master thesis, University of South Bohemia, České Budějovice.

Trophic mode

Parasitism and mycoheterotrophy: autotrophic

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Parasitism and mycoheterotrophy
Plant parasitism is based on either of two mechanisms. The first group of parasitic plants involves those parasitizing directly on another plant. These plants are called haustorial parasites. They take resources from the host’s vascular bundles using a specialized organ, the haustorium. The second group comprises mycoheterotrophic plants, which parasitize fungi via mycorrhizal interaction and gain organic carbon from them.

Plants in both groups display variable dependence on their host organism. The haustorial parasites include two distinct functional groups: green hemiparasites and holoparasites. Green hemiparasites are partial parasites that retain photosynthetic ability but obtain all mineral resources and a part of the organic carbon from the host. Holoparasites are non-green full parasites unable to photosynthesize. Location of the haustorial attachment to the host (root or stem) is another essential functional trait. The distinction between partial and full parasitism in haustorial parasites may not be straightforward. In the Czech flora, it is nevertheless possible to distinguish between stem hemi- and holoparasites, which are difficult to separate on the global scale (Těšitel 2016). Consequently, we use a traditional classification here and classify as holoparasites those plants that are in adulthood mostly without chlorophyll, even though some of them might have some chlorophyll and perform residual photosynthesis (e.g. Cuscuta).

In mycoheterotrophic plants, there is a continuum from initial mycoheterotrophs through partial mycoheterotrophs to full mycoheterotrophs. In the initial mycoheterotrophs, only initial stages, i.e. gametophytes or seedlings, are dependent on the fungus, whereas adult plants are autotrophic, while still depending on mycorrhizal symbiosis as a source of water and mineral nutrients. In the partial mycoheterotrophs, photosynthesizing adults obtain from their mycorrhizal fungi not only water and mineral nutrients but also different amounts of organic carbon. The full mycoheterotrophs lost their chlorophyll and are thus fully parasitic. In some partial mycoheterotrophs (e.g. the genus Cephalanthera), chlorotic individuals can be found, which lack chlorophyll and fully depend on their hosts.

Classification of haustorial parasites follows Těšitel (2016) with a further distinction of stem hemi- and holoparasites, and identification of mycoheterotrophs follows Merckx (2012).

Categories

autotrophic

root hemiparasite

stem hemiparasite

root holoparasite

stem holoparasite

partial or initial mycoheterotroph

full mycoheterotroph

Data source and citation

Těšitel J., Těšitelová T., Blažek P. & Lepš J. (2016) Parasitism and mycoheterotrophy. – www.pladias.cz.

Further references

Těšitel J. (2016) Functional biology of parasitic plants: a review. – Plant Ecology and Evolution 149: 5–20.
Merckx V. S. F. T. (2012) Mycoheterotrophy: the biology of plants living on fungi. – Springer, Berlin.
Carnivory: non-carnivorous

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Carnivory
Carnivorous plants attract, trap and kill their prey, animals (mainly insects and small crustaceans) and protozoans, and subsequently absorb the nutrients from their dead bodies.

Categories

carnivorous

non-carnivorous

Data source and citation

Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds) (1988) Květena České socialistické republiky [Flora of the Czech Socialist Republic]. Vol. 1. – Academia, Praha.
Hejný S., Slavík B., Hrouda L. & Skalický V. (eds) (1990) Květena České republiky [Flora of the Czech Republic]. Vol. 2. – Academia, Praha.
Hejný S., Slavík B., Kirschner J. & Křísa B. (eds) (1992) Květena České republiky [Flora of the Czech Republic]. Vol. 3. – Academia, Praha.
Slavík B., Chrtek J. jun. & Štěpánková J. (eds) (2000) Květena České republiky [Flora of the Czech Republic]. Vol. 6. – Academia, Praha.
Slavík B., Chrtek J. jun. & Tomšovic P. (eds) (1997) Květena České republiky [Flora of the Czech Republic]. Vol. 5. – Academia, Praha.
Slavík B., Smejkal M., Dvořáková M. & Grulich V. (eds) (1995) Květena České republiky [Flora of the Czech Republic]. Vol. 4. – Academia, Praha.
Slavík B., Štěpánková J. & Štěpánek J. (eds) (2004) Květena České republiky [Flora of the Czech Republic]. Vol. 7. – Academia, Praha.
Štěpánková J., Chrtek J. jun. & Kaplan Z. (eds) (2010) Květena České republiky [Flora of the Czech Republic]. Vol. 8. – Academia, Praha.
Symbiotic nitrogen fixation: no nitrogen-fixing symbionts

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Symbiotic nitrogen fixation
Plants are classified into those without symbiotic nitrogen fixers and those that form a symbiosis with nitrogen-fixing bacteria. The latter are further divided into those forming a symbiosis with rhizobia (e.g. Allorhizobium, Bradyrhizobium, Mesorhizobium, Rhizobium and Sinorhizobium) and those forming the actinorhizal symbiosis with the genus Frankia, the latter called actinorhizal plants (Bond 1983, Pawlowski & Sprent 2007, Sprent 2008, Benson 2016).

In the Czech flora, the rhizobial group is represented by virtually all legumes (family Fabaceae). Exceptions are three non-native cultivated woody species (Cercis siliquastrum, Gleditsia triacanthos, Gymnocladus dioicus) that do not nodulate (Tedersoo et al. 2018), which is generally considered as evidence for the absence of symbiosis. However, some studies suggest that functional nitrogen-fixing symbiosis may exist even without visible nodules (Bryan et al. 1996). Roots of Gleditsia triacanthos were recorded to contain bacterial structures similar to those in nodules with rhizobia, as well as the presence of nitrogenase (Faria et al. 2002). These genera also contain genes probably related to nodule formation, although their exact function is unclear (Graves et al. 1999). Because convincing evidence of nitrogen fixation in these species is missing, we consider them non-nitrogen-fixing for the time being.

Symbiosis with rhizobia was found in several other families (Tedersoo et al. 2018). Of these, the Czech flora includes only casually introduced Tribulus terrestris (Zygophyllaceae), in which a parallel infection with cyanobacteria was described (Sabet 1946, Mahmood & Athar 1998).

The actinorhizal group is represented in the Czech flora mainly by alder species (Alnus spp.) and also by cultivated species in the family Elaeagnaceae – Elaeagnus spp. and Hippophaë rhamnoides (Bond 1983, Benson 2016).

Categories

symbiosis with rhizobia

symbiosis with Frankia

no nitrogen-fixing symbionts

Data source and citation

Blažek P. & Lepš J. (2016) Symbiotic nitrogen fixation. – www.pladias.cz.

Further references

Benson D. R. (2016) Frankia & actinorhizal plants. – https://frankia.mcb.uconn.edu/ (accessed on 1 Feb 2021).
Bond G. (1983) Taxonomy and distribution of non-legume nitrogen-fixing systems. – In: Gordon J. C. & Wheeler C. T. (eds), Biological nitrogen fixation in forests: foundations and applications, p. 55–87, Martinus Nijhoff/Dr W. Junk Publ., The Hague.
Bryan J. A., Berlyn G. P. & Gordon J. C. (1996) Toward a new concept of the evolution of symbiotic nitrogen fixation in the Leguminosae. – Plant and Soil 186: 151–159.
de Faria S. M., Olivares F. L. & Xavier R. P. (2002) Nodule-structure in the roots of Gleditsia spp. a non-nodulating legume genus. – In: Pedrosa F. O., Hungria M., Yates G. & Newton W. E. (eds), Nitrogen fixation: from molecules to crop productivity. Current plant science and biotechnology in agriculture, vol 38. Springer, Dordrecht, p. 337.
Graves W. R., Foster C. M., Rosin F. M. & Schrader J. A. (1999) Two early nodulation genes are not markers for the capacity of leguminous nursery crops to form root nodules. – Journal of Environmental Horticulture 17: 126–129.
Mahmood A. & Athar M. (1998) Cyanobacterial root nodules in Tribulus terrestris L. (Zygophyllaceae). – In: Malik K. A. & Sajjad Mirza M. & Ladha J. K. (eds), Nitrogen fixation with non-legumes, Springer, Dordrecht, p. 345–350.
Pawlowski K. & Sprent J. I. (2007) Comparison between actinorhizal and legume symbioses. – In: Pawlowski K. & Newton W. E. (eds), Nitrogen-fixing actinorhizal symbioses, Springer, Dordrecht, p. 261–288.
Sabet Y. S. (1946) Bacterial root nodules in the Zygophyllaceae. – Nature 157: 656–657.
Sprent J. I. (2008) Evolution and diversity of legume symbiosis. – In: Dilworth M. J., James E. K., Sprent J. I. & Newton W. E. (eds), Nitrogen-fixing leguminous symbioses, Springer, Dordrecht, p. 1–21.
Tedersoo L., Laanisto L., Rahimlou S., Toussaint A., Hallikma T. & Pärtel M. (2018) Global database of plants with root-symbiotic nitrogen fixation: NodDB. – Journal of Vegetation Science 29: 560–568.

Taxon origin

Origin in the Czech Republic: native

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Origin in the Czech Republic
Taxa are classified according to whether they are native or alien to the Czech Republic. Following the definitions used in invasion ecology, native taxa are those that have evolved in the area of the Czech Republic or immigrated there without human assistance from the area where they had evolved. Alien taxa are those whose presence is a result of intentional or unintentional introduction by human activity and can be divided based on their residence time. The alien taxa are divided based on their residence time into archaeophytes and neophytes. Archaeophytes are taxa occurring in the wild that were introduced between the beginning of Neolithic agriculture and the year 1500, i.e. the beginning of intercontinental overseas trade after the discovery of the Americas. Neophytes are taxa occurring in the wild that were introduced after 1500 (see Richardson et al. 2000 for detailed definitions). Some taxa introduced in the Late Middle Ages or Early Modern Period, but with no exact information on the introduction date, were assigned to a joint category of Archaeophyte/neophyte. Additionally, some frequently cultivated taxa that are not known to have escaped from cultivation are listed as a separate category Cultivated. Category Lack of evidence of occurrence in the wild includes taxa for which spontaneous occurrence in the wild is doubtful. Taxa assigned to the category Absent in Czechia are not sufficiently supported by reliable records or occurred just once and disappeared.

The data included in the database follow the third edition of the Catalogue of alien plants of the Czech Republic (Pyšek et al. 2022 and references related to individual taxa therein).

Categories

native

archaeophyte

neophyte

archaeophyte/neophyte

cultivated only

lack of evidence of occurrence in the wild

absent in Czechia

Data source and citation

Pyšek P., Sádlo J., Chrtek J. Jr., Chytrý M., Kaplan Z., Pergl J., Pokorná A., Axmanová I., Čuda J., Doležal J., Dřevojan P., Hejda M., Kočár P., Kortz A., Lososová Z., Lustyk P., Skálová H., Štajerová K., Večeřa M., Vítková M., Wild J. & Danihelka J. (2022) Catalogue of alien plants of the Czech Republic (3rd edition): species richness, status, distributions, habitats, regional invasion levels, introduction pathways and impacts. – Preslia 94: 447–577.

Further references

Richardson D. M., Pyšek P., Rejmánek M., Barbour M. G., Panetta F. D. & West C. J. (2000) Naturalization and invasion of alien plants: concepts and definitions. – Diversity and Distributions 6: 93–107.

Ecological indicator values

Ellenberg-type indicator values

Light indicator value: 7 – half-light plant, mostly occurring at full light, but also in the shade up to about 30% of diffuse radiation incident in an open area

?

Temperature indicator value: 5x – moderate heat indicator, occurring from lowland to montane belt, mainly in submontane-temperate areas (generalist)

?

Moisture indicator value: 5x – indicator of fresh soils, focus on soils of average moisture, missing on wet and on soils that frequently dry out (generalist)

?

Reaction indicator value: 4x – transition between values 3 and 5 (generalist)

?

Nutrient indicator value: 3x – occurring at nutrient-poor sites more frequently than at average sites and exceptionally at rich sites (generalist)

?

Salinity indicator value: 0 – not salt tolerant, glycophyte

?

Habitat and sociology

Occurrence in habitats
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Occurrence in habitats

Data on taxon occurrence in habitats of the Czech Republic are based on the analysis of vegetation plots from the Czech National Phytosociological Database (Chytrý & Rafajová 2003) and its expert revision and completion based on the literature and field experience, especially for rare and taxonomically problematic taxa. The classification recognizes 88 basic habitats aggregated to 13 broader habitats that are defined by Sádlo et al. (2007: their Appendix 1):
1 Vegetation of cliffs, screes and walls
1A Calcareous cliffs
1B Siliceous cliffs and block fields
1C Walls
1D Mobile calcareous screes
2 Alpine and subalpine grasslands
2A Alpine grasslands on siliceous bedrock
2B Subalpine tall-forb and tall-grass vegetation
3 Aquatic vegetation
3A Macrophytic vegetation of eutrophic and mesotrophic still waters
3B Macrophytic vegetation of water streams
3C Macrophytic vegetation of oligotrophic lakes and pools
4 Wetland and riverine herbaceous vegetation
4A Reed-beds of eutrophic still waters
4B Halophilous reed and sedge beds
4C Eutrophic vegetation of muddy substrata
4D Riverine reed vegetation
4E Reed vegetation of brooks
4F Mesotrophic vegetation of muddy substrata
4G Tall-sedge beds
4H Vegetation of low annual hygrophilous herbs
4I Vegetation of nitrophilous annual hygrophilous herbs
4J River gravel banks
4K Petasites fringes of montane brooks
4L Nitrophilous herbaceous fringes of lowland rivers
5 Vegetation of springs and mires
5A Hard-water springs with tufa formation
5B Lowland to montane soft-water springs
5C Alpine and subalpine soft-water springs
5D Calcareous fens
5E Acidic moss-rich fens and peatland meadows
5F Transitional mires
5G Raised bogs
5H Wet peat soils and bog hollows
6 Meadows and mesic pastures
6A Mesic Arrhenatherum meadows
6B Montane mesic meadows
6C Pastures and park grasslands
6D Alluvial meadows of lowland rivers
6E Wet Cirsium meadows
6F Intermittently wet Molinia meadows
6G Vegetation of wet disturbed soils
7 Acidophilous grasslands
7A Subalpine and montane acidophilous grasslands
7B Submontane Nardus grasslands
8 Dry grasslands
8A Hercynian dry grasslands on rock outcrops
8B Submediterranean dry grasslands on rock outcrops
8C Narow-leaved sub-continental steppes
8D Broad-leaved dry grasslands
8E Acidophilous dry grasslands
8F Thermophilous forest fringe vegetation
9 Sand grasslands and rock-outcrop vegetation
9B Open vegetation of acidic sands
9C Festuca grasslands on acidic sands
9D Pannonian sand steppes
9E Acidophilous vegetation of spring therophytes and succulents
9F Basiphilous vegetation of spring therophytes and succulents
10 Saline vegetation
10G Continental vegetation of annual halophilous grasses
10H Inland vegetation of succulent halophytes
10I Inland saline meadows
10J Saline steppes
11 Heathlands and scrub
11A Dry lowland to subalpine heathlands
11D Subalpine acidophilous Pinus mugo scrub
11H Subalpine deciduous scrub
11I Willow carrs
11J Willow galleries of loamy and sandy river banks
11L Tall mesic and xeric shrub
11N Low xeric scrub
11R Scrub and pioneer woodland of forests clearings
12 Forests
12A Alder carrs
12B Alluvial forests
12C Oak-hornbeam forests
12D Ravine forests
12E Herb-rich beech forests
12F Limestone beech forests
12G Acidophilous beech forests
12H Peri-Alpidic basiphilous thermophilous oak forests
12I Sub-continental thermophilous oak forests
12J Acidophilous thermophilous oak forests
12K Acidophilous oak forests
12L Boreo-continental pine forests
12O Peri-Alpidic pine forests
12P Peatland pine forests
12Q Peatland birch forests
12R Acidophilous spruce forests
12S Basiphilous spruce forests
12T Robinia pseudoacacia plantations
12U Plantations of broad-leaved non-native trees
12V Picea plantations
12W Pinus and Larix plantations
13 Anthropogenic vegetation
13A Annual vegetation of ruderal habitats
13B Annual vegetation of arable land
13C Annual vegetation of trampled habitats
13D Perennial thermophilous ruderal vegetation
13E Perennial nitrophilous herbaceous vegetation of mesic sites
13F Herbaceous vegetation of forests clearings and Rubus scrub
Taxon occurrence in each habitat is assessed on a four-degree scale:
1 – occurrence – the taxon can grow in the habitat, but it tends to be rare there, and the habitat is not its ecological optimum
2 – optimum – the habitat or a part of it is the ecological optimum for this taxon
3 – dominant – the taxon can be assigned to the previous category and at the same time it frequently attains a cover above 25% in areas of 10–100 m² or 100–1000 m² in herbaceous or woody vegetation, respectively
4 – constant dominant – same as for the previous category but the taxon also determines the general appearance of the habitat (e.g. Calluna vulgaris in heathlands), occurring in ≥ 40% of the localities of the habitat

Data source and citation

Sádlo J., Chytrý M. & Pyšek P. (2007) Regional species pools of vascular plants in habitats of the Czech Republic. – Preslia 79: 303–321.

Further references

Chytrý M. & Rafajová M. (2003) Czech National Phytosociological Database: basic statistics of the available vegetation-plot data. – Preslia 75: 1–15.

5 Vegetation of springs and mires

5E Acidic moss-rich fens and peatland meadows: 1 – rare occurrence

6 Meadows and mesic pastures

6C Pastures and park grasslands: 1 – rare occurrence

6E Wet Cirsium meadows: 1 – rare occurrence

6F Intermittently wet Molinia meadows: 1 – rare occurrence

7 Acidophilous grasslands

7B Submontane Nardus grasslands: 1 – rare occurrence

11 Heathlands and scrub

11I Willow carrs: 1 – rare occurrence

11R Scrub and pioneer woodland of forests clearings: 2 – optimum

12 Forests

12A Alder carrs: 1 – rare occurrence

12B Alluvial forests: 1 – rare occurrence

12C Oak-hornbeam forests: 1 – rare occurrence

12K Acidophilous oak forests: 2 – optimum

12L Boreo-continental pine forests: 1 – rare occurrence

Affinity to the forest environment
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Affinity to the forest environment

The affinity of taxa to the forest environment is assessed using the categories of the German national list of forest taxa (Schmidt et al. 2011). Each taxon is assessed separately for the region of Thermophyticum (lowlands with thermophilous and drought-adapted flora) and merged regions of Mesophyticum and Oreophyticum (mid-elevations and mountains with mesophilous and mountain flora; Skalický 1988). The compilation was based on the list of regional species pools of Czech habitats (Sádlo et al. 2007), expert knowledge and various literature sources. It has been integrated into the European forest plant species list (Heinken et al. 2019).

Categories

0 – taxon does not spontaneously occur in Czech forests

1.1 – taxon occurring mainly in closed forests

1.2 – taxon occurring mainly along forest edges and in forest openings, including forest roads and paths, windthrow sites, burnt sites and forest clearings

2.1 – taxon occurring both in forest and open vegetation

2.2 – taxon occurring partly in forest but mainly in open vegetation

Data source and citation

Dřevojan P., Chytrý M., Sádlo J. & Pyšek P. (2016) Affinity to the forest environment. – www.pladias.cz.

Further references

Heinken T., Diekmann M., Liira J., Orczewska A., Brunet J., Chytrý M., Chabrerie O., de Frenne P., Decoq G., Dřevojan P., Dzwonko Z., Ewald J., Feilberg J., Graae B. J., Grytnes J. A., Hermy M., Kriebitzsch W.-U., Laivins M., Lindmo S., Marage D., Marozas V., Meirland A., Niemeyer T., Paal J., Pyšek P., Roosaluste E., Sádlo J., Schaminée J., Schmidt M., Tyler T., Verheyen K. & Wulf M. (2019) European forest plant species list. – Fighshare, https://doi.org/10.6084/m9.figshare.8095217.v1.
Sádlo J., Chytrý M. & Pyšek P. (2007) Regional species pools of vascular plants in habitats of the Czech Republic. – Preslia 79: 303–321.
Schmidt M., KriebitzschW.-U. & Ewald J. (eds) (2011) Waldartenlisten der Farn- und Blütenpflanzen, Moose und Flechten Deutschlands. – BfN-Skripten 299: 1–111.
Skalický V. (1988) Regionálně fytogeografické členění [Regional phytogeographic division]. – In: Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds), Květena České socialistické republiky [Flora of the Czech Socialist Republic] 1: 103–121, Academia, Praha.

Affinity to the forest environment in Thermophyticum: 2.1 – taxon occurring both in the forest and open vegetation

Affinity to the forest environment in Mesophyticum and Oreophyticum: 2.1 – taxon occurring both in the forest and open vegetation

Distribution and frequency

Elevational belt in the Czech Republic: submontane belt, montane belt

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Elevational belt in the Czech Republic
The lowest and the highest elevational vegetation belt in which the taxon commonly occurs in the Czech Republic. For some taxa, also extremes are shown, i.e. elevational belts in which the taxon rarely occurs outside its main elevational range. The submontane belt comprises merged supracolline and submontane belts, and the montane belt comprises merged montane and supramontane belts according to the classification of elevational vegetation belts used in the Flora of the Czech Republic (Skalický 1988). The data were taken from the Key to the Flora of the Czech Republic (Kaplan et al. 2019).

Categories

lowlands

colline belt

submontane belt

montane belt

subalpine belt

Data source and citation

Kaplan Z., Danihelka J., Chrtek J. Jr., Kirschner J., Kubát K., Štěpánek J. & Štech M. (eds) (2019) Klíč ke květeně České republiky [Key to the flora of the Czech Republic]. Ed. 2. – Academia, Praha.

Further references

Skalický V. (1988) Regionálně fytogeografické členění [Regional phytogeographic division]. – In: Hejný S., Slavík B., Chrtek J., Tomšovic P. & Kovanda M. (eds), Květena České socialistické republiky [Flora of the Czech Socialist Republic] 1: 103–121, Academia, Praha.
Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells: 27, 41

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Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells

The number of basic grid mapping cells (Central European Basic Area, CEBA) and the number of quadrants of the Central European flora mapping in that the taxon has been recorded within the territory of the Czech Republic are generated dynamically from the current occurrence records in the species distribution module of the Pladias Database. The basic grid cells measure 10 minutes in the west–east direction and 6 minutes in the south–north direction, which corresponds to approximately 12.0 × 11.1 km (133.2 km²) on the 50th parallel. The Czech Republic comprises 679 basic cells, including incomplete cells on the state borders. The quadrants are the basic grid cells divided into four. They measure 5 minutes in the west–east direction and 3 minutes in the south–north direction, which corresponds to approximately 6.0 × 5.55 km (33.3 km²) on the 50th parallel. Revised occurrence records marked as erroneous or uncertain are not counted.

Data source and citation

Pladias. Database of the Czech flora and vegetation. – www.pladias.cz.
Number of habitats with taxon occurrence in the Czech Republic
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Number of habitats with taxon occurrence in the Czech Republic

The number of habitat types (habitats) in which the taxon occurs was counted based on the data from the Czech National Phytosociological Database (Chytrý & Rafajová 2003) and their expert revision and completion, especially for rare and taxonomically problematic taxa. This number is a measure of the taxon’s ecological range. The classification recognizes 88 basic habitats aggregated into 13 broader habitats that are defined by Sádlo et al. (2007: their Appendix 1). The number of habitats is defined in four ways:

Number of narrow habitats in which the taxon occurs – the number of habitats of the total number of 88 in which the taxon occurs; the taxon may or may not have its ecological optimum in these habitats

Number of narrow habitats in which the taxon has its optimum – the number of habitats of the total number of 88 in which the taxon occurs and at the same time has its ecological optimum there (it may also be a dominant or constant dominant)

Number of broad habitats in which the taxon occurs – the number of habitats of the total number of 13 in which the taxon occurs; the taxon may but does not necessarily have its ecological optimum in these habitats

Number of broad habitats in which the taxon has its optimum – the number of habitats of the total number of 13 in which the taxon occurs and at the same time has its ecological optimum there (it may also be a dominant or constant dominant)

Data source and citation

Sádlo J., Chytrý M. & Pyšek P. (2007) Regional species pools of vascular plants in habitats of the Czech Republic. – Preslia 79: 303–321.

Further references

Chytrý M. & Rafajová M. (2003) Czech National Phytosociological Database: basic statistics of the available vegetation-plot data. – Preslia 75: 1–15.

Number of narrow habitats in which the taxon occurs: 12

Number of narrow habitats in which the taxon has its optimum: 2

Number of broad habitats in which the taxon occurs: 5

Number of broad habitats in which the taxon has its optimum: 2

Threats and protection

Red List 2017 (national categories): taxon is not on the Red List

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Red List 2017 (national categories)
National Red List categories were taken from the 2017 edition of the Red List of Vascular Plants of the Czech Republic (Grulich 2017). These categories, introduced in the previous editions of the Czech Red List, are different from the IUCN Red List categories. The main category “A” includes extinct or missing taxa, while the main category “C” includes endangered, near threatened and data deficient taxa.

Categories

A1 – extinct taxon

A2 – missing taxon

A3 – extinct or missing taxon (uncertain case)

C1r – critically threatened taxon, rare

C1t – critically threatened taxon, declining

C1b – critically threatened taxon, rare and declining

C2r – endangered taxon, rare

C2t – endangered taxon, declining

C2b – endangered taxon, rare and declining

C3 – vulnerable taxon

C4a – near threatened taxon

C4b – data deficient taxon

taxon is not on the Red List

Data source and citation

Grulich V. (2017) Červený seznam cévnatých rostlin ČR [The Red List of vascular plants of the Czech Republic]. – Příroda 35: 75–132.
Red List 2017 (IUCN categories): LC(NA) – least concern (taxon is not on the Red List)

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Red List 2017 (IUCN categories)
International Red List categories defined by the IUCN were taken from the 2017 edition of the Red List of Vascular Plants of the Czech Republic (Grulich 2017). Taxon assignments to these categories follow the internationally accepted rules (IUCN 2012, 2014). To some extent, the definitions of these categories differ from the national categories used in the previous Czech Red Lists. The national Red List included only threatened or possibly threatened taxa, implying that the non-included taxa are not threatened. Therefore, the non-included taxa are classified here as LC(NA) – least concern (taxon is not on the Red List).

Categories

EX – extinct

RE – regionally extinct

CR – critically endangered

EN – endangered

VU – vulnerable

NT – near threatened

LC – least concern

LC(NA) – least concern (taxon is not on the Red List)

DD – data deficient

NA – not applicable

NE – not evaluated

Data source and citation

Grulich V. (2017) Červený seznam cévnatých rostlin ČR [The Red List of vascular plants of the Czech Republic]. – Příroda 35: 75–132.

Further references

IUCN (2012) Guidelines for application of IUCN Red List criteria at regional and national levels. Version 4.0. – IUCN, Gland.
IUCN (2014) Guidelines for using the IUCN Red List categories and criteria. Version 11. – IUCN, Gland.
Legal protection: not protected by law

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Legal protection
Legal protection in the Czech Republic concerns the specifically protected species, i.e. rare taxa, threatened taxa and taxa significant from a cultural or scientific point of view that are listed in Annex II of the Decree of the Ministry of the Environment no. 395/1992. They comprise 487 taxa of vascular plants divided into three categories according to their vulnerability: critically threatened, endangered and vulnerable.

Categories

critically threatened taxon

endangered taxon

vulnerable taxon

not protected by law

Data source and citation

Decree no. 395/1992 of the Ministry of the Environment of the Czech Republic.

Leaf

Flower

Fruit, seed and dispersal

Trophic mode

Taxon origin

Ecological indicator values

Ellenberg-type indicator values

Habitat and sociology

Occurrence in habitats

5 Vegetation of springs and mires

6 Meadows and mesic pastures

7 Acidophilous grasslands

11 Heathlands and scrub

12 Forests

Affinity to the forest environment

Distribution and frequency

Number of habitats with taxon occurrence in the Czech Republic

Threats and protection