Galium pumilum agg.

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Habitus and growth type

  • Height [m]: 0.07–0.5
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  • Growth form: polycarpic perennial non-clonal herb
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  • Life form: hemicryptophyte
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  • Life strategy: CSR – competitor/stress-tolerator/ruderal
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Leaf

  • Leaf presence and metamorphosis: leaves present, not modified
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  • Leaf arrangement (phyllotaxis): verticillate
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  • Leaf shape: simple – entire
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  • Stipules: present
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  • Petiole: absent
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  • Leaf life span: summer green
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  • Leaf anatomy: mesomorphic
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Flower

  • Flowering period [month]: May–July
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  • Flowering phase: 6 Cornus sanguinea-Melica uniflora (start of early summer)
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  • Flower colour: white
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  • Flower symmetry: actinomorphic
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  • Perianth type: calyx absent, corolla present
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  • Perianth fusion: fused
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  • Shape of the sympetalous corolla or syntepalous perianth: rotate
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  • Inflorescence type: panicula e dichasiis composita
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  • Dicliny: synoecious
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  • Generative reproduction type: mixed mating
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  • Pollination syndrome: insect-pollination, selfing
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Fruit, seed and dispersal

  • Fruit type: dry fruit – pair of nutlets
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  • Fruit colour: brown, black
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  • Reproduction type: by seed/spores and vegetatively
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  • Dispersal unit (diaspore): fruit, infrutescence or its part
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  • Dispersal strategy: Allium (mainly autochory)
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  • Myrmecochory: non-myrmecochorous (b)
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Belowground organs and clonality

  • Shoot metamorphosis: stolon, pleiocorm
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  • Storage organ: stolon, pleiocorm
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  • Shoot life span (cyclicity): monocyclic shoots prevailing
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  • Branching type of stem-derived organs of clonal growth: sympodial
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  • Primary root: present
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  • Bud bank

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  • Number of buds per shoot at the soil surface (root buds excluded):
  • Number of buds per shoot at a depth of 0–10 cm (root buds excluded):
  • Number of buds per shoot at a depth greater than 10 cm (root buds excluded):
  • Size of the belowground bud bank (root buds excluded):
  • Depth of the belowground bud bank (root buds exluded) [cm]:
  • Number of buds per shoot at the soil surface (root buds included):
  • Number of buds per shoot at a depth of 0–10 cm (root buds included):
  • Number of buds per shoot at a depth greater than 10 cm (root buds included):
  • Size of the belowground bud bank (root buds included):
  • Depth of the belowground bud bank (root buds included) [cm]:

Trophic mode

  • Parasitism and mycoheterotrophy: autotrophic
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  • Carnivory: non-carnivorous
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  • Symbiotic nitrogen fixation: no nitrogen-fixing symbionts
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Karyology

  • Chromosome number (2n): 22, 44
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  • Ploidy level (x): 2, 4
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  • 2C genome size [Mbp]: 1409.25
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  • 1Cx monoploid genome size [Mbp]: 424.62
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Taxon origin

  • Origin in the Czech Republic: native
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Ecological indicator values

  • Ellenberg-type indicator values

  • Light indicator value: 7 – half-light plant, mostly occurring at full light, but also in the shade up to about 30% of diffuse radiation incident in an open area
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  • Temperature indicator value: 5 – moderate heat indicator, occurring from lowland to montane belt, mainly in submontane-temperate areas
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  • Moisture indicator value: 4 – transition between values 3 and 5
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  • Reaction indicator value: 5 – indicator of moderate acidity, occurring rarely in strongly acidic as well as in neutral to alkaline conditions
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  • Nutrient indicator value: 3 – occurring at nutrient-poor sites more frequently than at average sites and exceptionally at rich sites
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  • Salinity indicator value: 0 – not salt tolerant, glycophyte
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  • Indicator values for disturbance

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  • Whole-community disturbance frequency indicator value: -1.42
  • Herb layer disturbance frequency indicator value: -0.51
  • Whole-community disturbance severity indicator value: 0.26
  • Herb layer disturbance severity indicator value: 0.24
  • Whole-community structure based disturbance indicator value: 0.33
  • Herb layer structure-based disturbance indicator value: 0.49

Habitat and sociology

  • Occurrence in habitats

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  • 1 Vegetation of cliffs, screes and walls

  • 1A Calcareous cliffs: 1 – rare occurrence
  • 1B Siliceous cliffs and block fields: 2 – optimum
  • 1D Mobile calcareous screes: 1 – rare occurrence
  • 2 Alpine and subalpine grasslands

  • 2B Subalpine tall-forb and tall-grass vegetation: 1 – rare occurrence
  • 5 Vegetation of springs and mires

  • 5E Acidic moss-rich fens and peatland meadows: 1 – rare occurrence
  • 5F Transitional mires: 1 – rare occurrence
  • 6 Meadows and mesic pastures

  • 6A Mesic Arrhenatherum meadows: 2 – optimum
  • 6B Montane mesic meadows: 2 – optimum
  • 6C Pastures and park grasslands: 2 – optimum
  • 6D Alluvial meadows of lowland rivers: 1 – rare occurrence
  • 6E Wet Cirsium meadows: 1 – rare occurrence
  • 6F Intermittently wet Molinia meadows: 1 – rare occurrence
  • 7 Acidophilous grasslands

  • 7A Subalpine and montane acidophilous grasslands: 1 – rare occurrence
  • 7B Submontane Nardus grasslands: 2 – optimum
  • 8 Dry grasslands

  • 8A Hercynian dry grasslands on rock outcrops: 2 – optimum
  • 8B Submediterranean dry grasslands on rock outcrops: 2 – optimum
  • 8C Narrow-leaved sub-continental steppes: 2 – optimum
  • 8D Broad-leaved dry grasslands: 2 – optimum
  • 8E Acidophilous dry grasslands: 2 – optimum
  • 8F Thermophilous forest fringe vegetation: 2 – optimum
  • 9 Sand grasslands and rock-outcrop vegetation

  • 9C Festuca grasslands on acidic sands: 1 – rare occurrence
  • 9E Acidophilous vegetation of spring therophytes and succulents: 1 – rare occurrence
  • 9F Basiphilous vegetation of spring therophytes and succulents: 1 – rare occurrence
  • 11 Heathlands and scrub

  • 11A Dry lowland to subalpine heathlands: 2 – optimum
  • 11N Low xeric scrub: 1 – rare occurrence
  • 11R Scrub and pioneer woodland of forests clearings: 1 – rare occurrence
  • 12 Forests

  • 12C Oak-hornbeam forests: 1 – rare occurrence
  • 12D Ravine forests: 1 – rare occurrence
  • 12F Limestone beech forests: 2 – optimum
  • 12G Acidophilous beech forests: 1 – rare occurrence
  • 12H Peri-Alpidic basiphilous thermophilous oak forests: 2 – optimum
  • 12I Sub-continental thermophilous oak forests: 2 – optimum
  • 12J Acidophilous thermophilous oak forests: 2 – optimum
  • 12K Acidophilous oak forests: 2 – optimum
  • 12L Boreo-continental pine forests: 2 – optimum
  • 12O Peri-Alpidic pine forests: 2 – optimum
  • 12T Robinia pseudacacia plantations: 1 – rare occurrence
  • 12V Spruce plantations: 1 – rare occurrence
  • 12W Pine and larch plantations: 2 – optimum
  • Ecological specialization indices

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  • Ecological specialization index for all vegetation types: 4.4
  • Ecological specialization index for non-forest vegetation: 5
  • Ecological specialization index for forest vegetation: 4.7

Distribution and frequency

  • Floristic zone: northern temperate, southern temperate, submeridional
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  • Floristic region: Europe
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  • Elevational belt in the Czech Republic: lowlands, colline belt, submontane belt, montane belt, subalpine belt
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  • Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells: 579, 1544
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  • Commonness in vegetation plots from the Czech Republic

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  • Occurrence frequency in vegetation plots: 1.6 %
  • Occurrence frequency in vegetation plots with a cover above 5%: 1.9 %
  • Occurrence frequency in vegetation plots with a cover above 25%: 0.4 %
  • Occurrence frequency in vegetation plots with a cover above 50%: 0 %
  • Mean percentage cover in vegetation plots: 2.3 %
  • Maximum percentage cover in vegetation plots: 38 %
  • Number of habitats with taxon occurrence in the Czech Republic

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  • Number of narrow habitats in which the taxon occurs: 39
  • Number of narrow habitats in which the taxon has its optimum: 20
  • Number of broad habitats in which the taxon occurs: 9
  • Number of broad habitats in which the taxon has its optimum: 6
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