Gymnocarpium robertianum

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Habitus and growth type

  • Height [m]: 0.15–0.5
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  • Growth form: clonal herb
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  • Life form: geophyte
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  • Life strategy: CS – competitor/stress-tolerator
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Leaf

  • Leaf presence and metamorphosis: leaves present, not modified
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  • Leaf arrangement (phyllotaxis): alternate
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  • Leaf shape: compound – bipinnate, compound – tripinnate
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  • Stipules: absent
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  • Petiole: present
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  • Leaf life span: summer green
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  • Leaf anatomy: hygromorphic
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Flower

  • Flowering period [month]: June–August
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  • Dicliny: synoecious
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Fruit, seed and dispersal

  • Reproduction type: by seed/spores and vegetatively
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  • Dispersal unit (diaspore): seed
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  • Dispersal strategy: Lycopodium (mainly anemochory)
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  • Myrmecochory: non-myrmecochorous (b)
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Belowground organs and clonality

  • Shoot metamorphosis: rhizome
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  • Storage organ: rhizome
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  • Type of clonal growth organ: hypogeogenous rhizome
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  • Freely dispersible organs of clonal growth: absent
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  • Shoot life span (cyclicity): monocyclic shoots prevailing
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  • Branching type of stem-derived organs of clonal growth: dichotomous
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  • Primary root: absent
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  • Persistence of the clonal growth organ [year]:
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  • Number of clonal offspring: 0.5
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  • Lateral spreading distance by clonal growth [m]: 0.13
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  • Clonal index: 3
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  • Bud bank

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  • Number of buds per shoot at the soil surface (root buds excluded):
  • Number of buds per shoot at a depth of 0–10 cm (root buds excluded):
  • Number of buds per shoot at a depth greater than 10 cm (root buds excluded):
  • Size of the belowground bud bank (root buds excluded):
  • Depth of the belowground bud bank (root buds exluded) [cm]:
  • Number of buds per shoot at the soil surface (root buds included):
  • Number of buds per shoot at a depth of 0–10 cm (root buds included):
  • Number of buds per shoot at a depth greater than 10 cm (root buds included):
  • Size of the belowground bud bank (root buds included):
  • Depth of the belowground bud bank (root buds included) [cm]:

Trophic mode

  • Parasitism and mycoheterotrophy: autotrophic
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  • Carnivory: non-carnivorous
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  • Symbiotic nitrogen fixation: no nitrogen-fixing symbionts
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Karyology

  • Chromosome number (2n): 160
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  • Ploidy level (x): 4
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  • 2C genome size [Mbp]: 11124.72
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  • 1Cx monoploid genome size [Mbp]: 2781.18
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  • Genomic GC content: 43.6 %
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Taxon origin

  • Origin in the Czech Republic: native
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Ecological indicator values

  • Ellenberg-type indicator values

  • Light indicator value: 7 – half-light plant, mostly occurring at full light, but also in the shade up to about 30% of diffuse radiation incident in an open area
    ?
  • Temperature indicator value: 5 – moderate heat indicator, occurring from lowland to montane belt, mainly in submontane-temperate areas
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  • Moisture indicator value: 5 – indicator of fresh soils, focus on soils of average moisture, missing on wet and on soils that frequently dry out
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  • Reaction indicator value: 8 – transition between values 7 and 9, occurring mostly in calcium-rich conditions
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  • Nutrient indicator value: 4 – transition between values 3 and 5
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  • Salinity indicator value: 0 – not salt tolerant, glycophyte
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  • Indicator values for disturbance

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  • Whole-community disturbance frequency indicator value: -1.38
  • Herb layer disturbance frequency indicator value: -1.14
  • Whole-community disturbance severity indicator value: 0.19
  • Herb layer disturbance severity indicator value: 0.2
  • Whole-community structure based disturbance indicator value: 0.49
  • Herb layer structure-based disturbance indicator value: 0.26

Habitat and sociology

  • Occurrence in habitats

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  • 1 Vegetation of cliffs, screes and walls

  • 1A Calcareous cliffs: 2 – optimum
  • 1B Siliceous cliffs and block fields: 1 – rare occurrence
  • 1C Walls: 1 – rare occurrence
  • 1D Mobile calcareous screes: 2 – optimum
  • 5 Vegetation of springs and mires

  • 5B Lowland to montane soft-water springs: 1 – rare occurrence
  • 8 Dry grasslands

  • 8B Submediterranean dry grasslands on rock outcrops: 1 – rare occurrence
  • 11 Heathlands and scrub

  • 11L Tall mesic and xeric shrub: 1 – rare occurrence
  • 12 Forests

  • 12D Ravine forests: 1 – rare occurrence
  • 12E Herb-rich beech forests: 1 – rare occurrence
  • 12F Limestone beech forests: 1 – rare occurrence
  • 12O Peri-Alpidic pine forests: 1 – rare occurrence
  • Affinity to the forest environment

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  • Affinity to the forest environment in Thermophyticum: 2.1 – taxon occurring both in the forest and open vegetation
  • Affinity to the forest environment in Mesophyticum and Oreophyticum: 2.1 – taxon occurring both in the forest and open vegetation
  • Diagnostic taxon

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  • Diagnostic taxon of classes: SC Thlaspietea rotundifolii
  • Diagnostic taxon of alliances: SAA Cystopteridion, SCA Stipion calamagrostis
  • Diagnostic taxon of associations: SCA01 Gymnocarpietum robertiani
  • Constant taxon

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  • Constant taxon of associations: SCA01 Gymnocarpietum robertiani
  • Dominant taxon

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  • Dominant taxon of associations: SCA01 Gymnocarpietum robertiani
  • Ecological specialization indices

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  • Ecological specialization index for all vegetation types: 3.7
  • Ecological specialization index for non-forest vegetation: 4
  • Ecological specialization index for forest vegetation: 5.3

Distribution and frequency

  • Floristic zone: boreal, northern temperate, southern temperate, submeridional, meridional
    ?
  • Floristic region: circumpolar
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  • Continentality degree: 6
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  • Distribution range extension along the continentality gradient: 6
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  • Elevational belt in the Czech Republic: colline belt, submontane belt
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  • Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells: 313, 464
    ?
  • Commonness in vegetation plots from the Czech Republic

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  • Occurrence frequency in vegetation plots: 0.1 %
  • Occurrence frequency in vegetation plots with a cover above 5%: 70.4 %
  • Occurrence frequency in vegetation plots with a cover above 25%: 29.6 %
  • Occurrence frequency in vegetation plots with a cover above 50%: 11.1 %
  • Mean percentage cover in vegetation plots: 20.4 %
  • Maximum percentage cover in vegetation plots: 63 %
  • Number of habitats with taxon occurrence in the Czech Republic

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  • Number of narrow habitats in which the taxon occurs: 11
  • Number of narrow habitats in which the taxon has its optimum: 2
  • Number of broad habitats in which the taxon occurs: 5
  • Number of broad habitats in which the taxon has its optimum: 1

Threats and protection

  • Red List 2017 (national categories): taxon is not on the Red List
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  • Red List 2017 (IUCN categories): LC(NA) – least concern (taxon is not on the Red List)
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  • Legal protection: not protected by law
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