Marrubium vulgare

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Habitus and growth type

  • Height [m]: 0.3–0.7
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  • Growth form: polycarpic perennial non-clonal herb
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  • Life form: hemicryptophyte (chamaephyte)
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  • Life strategy: CSR – competitor/stress-tolerator/ruderal
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  • Life strategy (Pierce method based on leaf traits): CSR
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  • Life strategy (Pierce method, C-score): 33.7 %
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  • Life strategy (Pierce method, S-score): 40.8 %
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  • Life strategy (Pierce method, R-score): 25.5 %
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Leaf

  • Leaf presence and metamorphosis: leaves present, not modified
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  • Leaf arrangement (phyllotaxis): opposite
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  • Leaf shape: simple – entire
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  • Stipules: absent
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  • Petiole: present
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  • Leaf life span: summer green
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  • Leaf anatomy: scleromorphic, mesomorphic
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Flower

  • Flowering period [month]: July-September
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  • Flowering phase: 7 Ligustrum vulgare-Stachys sylvatica (end of early summer)
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  • Flower colour: white
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  • Flower symmetry: zygomorphic
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  • Perianth type: calyx and corolla
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  • Perianth fusion: fused
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  • Shape of the sympetalous corolla or syntepalous perianth: bilabiate
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  • Calyx fusion: synsepalous
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  • Inflorescence type: pseudospica e verticillastris composita
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  • Dicliny: synoecious, gynodioecious
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  • Generative reproduction type: mixed mating
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  • Pollination syndrome: insect-pollination, selfing
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Fruit, seed and dispersal

  • Fruit type: dry fruit – cluster of four one-seeded nutlets
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  • Fruit colour: brown
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  • Reproduction type: only by seed/spores
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  • Dispersal unit (diaspore): fruit, infrutescence or its part
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  • Dispersal strategy: Allium (mainly autochory)
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  • Myrmecochory: non-myrmecochorous (b)
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Belowground organs and clonality

  • Shoot metamorphosis: pleiocorm
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  • Storage organ: pleiocorm
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  • Shoot life span (cyclicity): monocyclic shoots prevailing
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  • Branching type of stem-derived organs of clonal growth: sympodial
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  • Primary root: present
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  • Bud bank

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  • Number of buds per shoot at the soil surface (root buds excluded): 5
  • Number of buds per shoot at a depth of 0–10 cm (root buds excluded): 5
  • Number of buds per shoot at a depth greater than 10 cm (root buds excluded): 0
  • Size of the belowground bud bank (root buds excluded): 10
  • Depth of the belowground bud bank (root buds exluded) [cm]: 3
  • Number of buds per shoot at the soil surface (root buds included): 5
  • Number of buds per shoot at a depth of 0–10 cm (root buds included): 5
  • Number of buds per shoot at a depth greater than 10 cm (root buds included): 0
  • Size of the belowground bud bank (root buds included): 10
  • Depth of the belowground bud bank (root buds included) [cm]: 3

Trophic mode

  • Parasitism and mycoheterotrophy: autotrophic
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  • Carnivory: non-carnivorous
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  • Symbiotic nitrogen fixation: no nitrogen-fixing symbionts
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Taxon origin

  • Origin in the Czech Republic: archaeophyte
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  • Invasion status: casual
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  • Geographic origin: Mediterranean
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  • Introduction pathway: accidental introduction
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Ecological indicator values

  • Ellenberg-type indicator values

  • Light indicator value: 9 – full light plant, occurring only in fully irradiated places, not at less than 50% of diffuse radiation incident in an open area
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  • Temperature indicator value: 7 – heat indicator, occurring in relatively warm lowlands
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  • Moisture indicator value: 4 – transition between values 3 and 5
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  • Reaction indicator value: 8 – transition between values 7 and 9, occurring mostly in calcium-rich conditions
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  • Nutrient indicator value: 8 – pronounced nutrient indicator
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  • Salinity indicator value: 1 – salt tolerant, mostly on low-salt to salt-free soils, but occasionally on slightly salty soils
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Habitat and sociology

  • Occurrence in habitats

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  • 11 Heathlands and scrub

  • 11R Scrub and pioneer woodland of forests clearings: 1 – rare occurrence
  • 12 Forests

  • 12T Robinia pseudacacia plantations: 1 – rare occurrence
  • 13 Anthropogenic vegetation

  • 13A Annual vegetation of ruderal habitats: 1 – rare occurrence
  • 13D Perennial thermophilous ruderal vegetation: 2 – optimum
  • Affinity to the forest environment

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  • Affinity to the forest environment in Thermophyticum: 0 – taxon that does not spontaneously occur in Czech forests
  • Affinity to the forest environment in Mesophyticum and Oreophyticum: 0 – taxon that does not spontaneously occur in Czech forests

Distribution and frequency

  • Floristic zone: northern temperate, southern temperate, submeridional, meridional
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  • Floristic region: Europe, Western Asia
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  • Distribution range extension along the continentality gradient: 6
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  • Elevational belt in the Czech Republic: lowlands, colline belt
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  • Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells: 185, 291
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  • Commonness in vegetation plots from the Czech Republic

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  • Occurrence frequency in vegetation plots: 0 %
  • Occurrence frequency in vegetation plots with a cover above 5%: 14.3 %
  • Occurrence frequency in vegetation plots with a cover above 25%: 0 %
  • Occurrence frequency in vegetation plots with a cover above 50%: 0 %
  • Mean percentage cover in vegetation plots: 3.6 %
  • Maximum percentage cover in vegetation plots: 13 %
  • Number of habitats with taxon occurrence in the Czech Republic

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  • Number of narrow habitats in which the taxon occurs: 4
  • Number of narrow habitats in which the taxon has its optimum: 1
  • Number of broad habitats in which the taxon occurs: 3
  • Number of broad habitats in which the taxon has its optimum: 1

Threats and protection

  • Red List 2017 (national categories): C1t – critically threatened taxon, declining
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  • Red List 2017 (IUCN categories): CR – critically endangered
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  • Legal protection: not protected by law
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