Pulsatilla grandis subsp. grandis

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Habitus and growth type

  • Height [m]: 0.05–0.4
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  • Growth form: polycarpic perennial non-clonal herb
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  • Life form: hemicryptophyte
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Leaf

  • Leaf presence and metamorphosis: leaves present, not modified
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  • Leaf arrangement (phyllotaxis): rosulate
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  • Leaf shape: simple – pinnately divided, compound – imparipinnate
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  • Stipules: absent
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  • Petiole: present
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Flower

  • Flowering period [month]: March–May
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  • Flower colour: violet
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  • Flower symmetry: actinomorphic
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  • Perianth type: homochlamydeous
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  • Perianth fusion: free
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  • Inflorescence type: flores solitarii
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  • Generative reproduction type: facultative allogamy
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Fruit, seed and dispersal

  • Fruit type: dry fruit – head of achenes
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  • Fruit colour: brown
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  • Dispersal unit (diaspore): fruit, infrutescence or its part
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  • Dispersal strategy: Epilobium (mainly anemochory and autochory)
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  • Myrmecochory: non-myrmecochorous (b)
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Belowground organs and clonality

  • Shoot life span (cyclicity): dicyclic or polycyclic shoots prevailing
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  • Branching type of stem-derived organs of clonal growth: sympodial
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  • Primary root: present
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  • Position of root buds: primary root
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  • Role of root buds in life-history of a plant: regenerative
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  • Bud bank

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  • Number of buds per shoot at the soil surface (root buds excluded): 5
  • Number of buds per shoot at a depth of 0–10 cm (root buds excluded): 15
  • Number of buds per shoot at a depth greater than 10 cm (root buds excluded): 0
  • Size of the belowground bud bank (root buds excluded): 20
  • Depth of the belowground bud bank (root buds exluded) [cm]: 4
  • Number of buds per shoot at the soil surface (root buds included): 5
  • Number of buds per shoot at a depth of 0–10 cm (root buds included): 15
  • Number of buds per shoot at a depth greater than 10 cm (root buds included): 0
  • Size of the belowground bud bank (root buds included): 20
  • Depth of the belowground bud bank (root buds included) [cm]: 4

Trophic mode

  • Parasitism and mycoheterotrophy: autotrophic
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  • Carnivory: non-carnivorous
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  • Symbiotic nitrogen fixation: no nitrogen-fixing symbionts
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Karyology

  • Chromosome number (2n): 32
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  • Ploidy level (x): 4
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  • 2C genome size [Mbp]: 26314.85
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  • 1Cx monoploid genome size [Mbp]: 6578.71
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  • Genomic GC content: 45.1 %
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Taxon origin

  • Origin in the Czech Republic: native
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Ecological indicator values

  • Ellenberg-type indicator values

  • Light indicator value: 8 – light plant, only exceptionally occurring at less than 40% of diffuse radiation incident in an open area
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  • Temperature indicator value: 7 – heat indicator, occurring in relatively warm lowlands
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  • Moisture indicator value: 3 – missing on damp soil
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  • Reaction indicator value: 7 – indicator of slightly acidic to slightly basic conditions, never occurring in very acidic conditions
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  • Nutrient indicator value: 2 – transition between values 1 and 3
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  • Salinity indicator value: 0 – not salt tolerant, glycophyte
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  • Indicator values for disturbance

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  • Whole-community disturbance frequency indicator value: -0.37
  • Herb layer disturbance frequency indicator value: -0.05
  • Whole-community disturbance severity indicator value: 0.2
  • Herb layer disturbance severity indicator value: 0.24
  • Whole-community structure based disturbance indicator value: 0.73
  • Herb layer structure-based disturbance indicator value: 0.76

Habitat and sociology

  • Occurrence in habitats

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  • 8 Dry grasslands

  • 8A Hercynian dry grasslands on rock outcrops: 1 – rare occurrence
  • 8B Submediterranean dry grasslands on rock outcrops: 2 – optimum
  • 8C Narrow-leaved sub-continental steppes: 2 – optimum
  • 8D Broad-leaved dry grasslands: 2 – optimum
  • 8E Acidophilous dry grasslands: 2 – optimum
  • 8F Thermophilous forest fringe vegetation: 1 – rare occurrence
  • 9 Sand grasslands and rock-outcrop vegetation

  • 9E Acidophilous vegetation of spring therophytes and succulents: 1 – rare occurrence
  • 9F Basiphilous vegetation of spring therophytes and succulents: 1 – rare occurrence
  • 11 Heathlands and scrub

  • 11A Dry lowland to subalpine heathlands: 1 – rare occurrence
  • 11N Low xeric scrub: 1 – rare occurrence
  • Affinity to the forest environment

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  • Affinity to the forest environment in Thermophyticum: 0 – taxon that does not spontaneously occur in Czech forests
  • Affinity to the forest environment in Mesophyticum and Oreophyticum: 0 – taxon that does not spontaneously occur in Czech forests
  • Diagnostic taxon

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  • Ecological specialization indices

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  • Ecological specialization index for all vegetation types: 5.7
  • Ecological specialization index for non-forest vegetation: 5.7
  • Colonization ability

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  • Index of colonization success (ICS): 2
  • Index of colonization potential (ICP): 2
  • Optimum successional age [years]: 75

Distribution and frequency

  • Continentality degree: 5
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  • Distribution range extension along the continentality gradient: 3
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  • Elevational belt in the Czech Republic: colline belt
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  • Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells: 71, 141
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  • Commonness in vegetation plots from the Czech Republic

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  • Occurrence frequency in vegetation plots: 0.3 %
  • Occurrence frequency in vegetation plots with a cover above 5%: 7.8 %
  • Occurrence frequency in vegetation plots with a cover above 25%: 0 %
  • Occurrence frequency in vegetation plots with a cover above 50%: 0 %
  • Mean percentage cover in vegetation plots: 3.1 %
  • Maximum percentage cover in vegetation plots: 13 %
  • Number of habitats with taxon occurrence in the Czech Republic

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  • Number of narrow habitats in which the taxon occurs: 10
  • Number of narrow habitats in which the taxon has its optimum: 4
  • Number of broad habitats in which the taxon occurs: 3
  • Number of broad habitats in which the taxon has its optimum: 1

Threats and protection

  • Red List 2017 (national categories): C2b – endangered taxon, rare and declining
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  • Red List 2017 (IUCN categories): VU – vulnerable
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  • Legal protection: endangered taxon
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