Dactylorhiza maculata agg.

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Habitus and growth type

  • Height [m]: 0.15
    ?
  • Growth form: clonal herb
    ?
  • Life form: geophyte
    ?
  • Life strategy: CSR – competitor/stress-tolerator/ruderal
    ?
  • Life strategy (Pierce method based on leaf traits): CR
    ?
  • Life strategy (Pierce method, C-score): 49.8 %
    ?
  • Life strategy (Pierce method, S-score): 0 %
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  • Life strategy (Pierce method, R-score): 50.2 %
    ?

Leaf

  • Leaf presence and metamorphosis: leaves present, not modified
    ?
  • Leaf arrangement (phyllotaxis): alternate
    ?
  • Leaf shape: simple – entire
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  • Stipules: absent
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  • Petiole: absent
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  • Leaf life span: summer green
    ?
  • Leaf anatomy: mesomorphic
    ?

Flower

  • Flowering period [month]: May–August
    ?
  • Flowering phase: 6 Cornus sanguinea-Melica uniflora (start of early summer)
    ?
  • Flower colour: white, pink, pink-violet, red-violet
    ?
  • Flower symmetry: zygomorphic
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  • Perianth type: homochlamydeous
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  • Perianth fusion: free
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  • Inflorescence type: spica
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  • Dicliny: synoecious
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  • Generative reproduction type: facultative allogamy
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  • Pollination syndrome: insect-pollination
    ?

Fruit, seed and dispersal

  • Fruit type: dry fruit – capsule
    ?
  • Fruit colour: green, brown
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  • Reproduction type: only by seed/spores
    ?
  • Dispersal unit (diaspore): seed
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  • Dispersal strategy: Lycopodium (mainly anemochory)
    ?
  • Myrmecochory: non-myrmecochorous (b)
    ?

Belowground organs and clonality

  • Root metamorphosis: root tuber
    ?
  • Storage organ: root tuber
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  • Type of clonal growth organ: root tuber
    ?
  • Freely dispersible organs of clonal growth: absent
    ?
  • Shoot life span (cyclicity): monocyclic shoots prevailing
    ?
  • Branching type of stem-derived organs of clonal growth: sympodial
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  • Primary root: absent
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  • Persistence of the clonal growth organ [year]:
    ?
  • Number of clonal offspring:
    ?
  • Lateral spreading distance by clonal growth [m]: 0.01
    ?
  • Clonal index: 3
    ?
  • Bud bank

    ?
  • Number of buds per shoot at the soil surface (root buds excluded): 1
  • Number of buds per shoot at a depth of 0–10 cm (root buds excluded):
  • Number of buds per shoot at a depth greater than 10 cm (root buds excluded):
  • Size of the belowground bud bank (root buds excluded): 6
  • Depth of the belowground bud bank (root buds exluded) [cm]: 4
  • Number of buds per shoot at the soil surface (root buds included): 1
  • Number of buds per shoot at a depth of 0–10 cm (root buds included):
  • Number of buds per shoot at a depth greater than 10 cm (root buds included):
  • Size of the belowground bud bank (root buds included): 6
  • Depth of the belowground bud bank (root buds included) [cm]: 4

Trophic mode

  • Parasitism and mycoheterotrophy: partial or initial mycoheterotroph
    ?
  • Carnivory: non-carnivorous
    ?
  • Symbiotic nitrogen fixation: no nitrogen-fixing symbionts
    ?

Karyology

  • Chromosome number (2n): 40
    ?
  • Ploidy level (x): 2
    ?
  • 2C genome size [Mbp]: 10595.65
    ?
  • 1Cx monoploid genome size [Mbp]: 5297.83
    ?

Taxon origin

  • Origin in the Czech Republic: native
    ?

Ecological indicator values

  • Ellenberg-type indicator values

  • Light indicator value: 6 – transition between values 5 and 7; rarely at less than 20% of diffuse radiation incident in an open area
    ?
  • Temperature indicator value: 4 – transition between values 3 and 5
    ?
  • Moisture indicator value: 7 – humidity indicator, focus on well moistened, but not wet soils
    ?
  • Reaction indicator value: 5x – indicator of moderate acidity, occurring rarely in strongly acidic as well as in neutral to alkaline conditions (generalist)
    ?
  • Nutrient indicator value: 3 – occurring at nutrient-poor sites more frequently than at average sites and exceptionally at rich sites
    ?
  • Salinity indicator value: 0 – not salt tolerant, glycophyte
    ?
  • Indicator values for disturbance

    ?
  • Whole-community disturbance frequency indicator value: -1.36
  • Herb layer disturbance frequency indicator value: -0.56
  • Whole-community disturbance severity indicator value: 0.2
  • Herb layer disturbance severity indicator value: 0.19
  • Whole-community structure based disturbance indicator value: 0.42
  • Herb layer structure-based disturbance indicator value: 0.55

Habitat and sociology

  • Occurrence in habitats

    ?
  • 2 Alpine and subalpine grasslands

  • 2B Subalpine tall-forb and tall-grass vegetation: 1 – rare occurrence
  • 4 Wetland and riverine herbaceous vegetation

  • 4K Petasites fringes of montane brooks: 1 – rare occurrence
  • 5 Vegetation of springs and mires

  • 5B Lowland to montane soft-water springs: 1 – rare occurrence
  • 5C Alpine and subalpine soft-water springs: 1 – rare occurrence
  • 5D Calcareous fens: 1 – rare occurrence
  • 5E Acidic moss-rich fens and peatland meadows: 2 – optimum
  • 5F Transitional mires: 1 – rare occurrence
  • 6 Meadows and mesic pastures

  • 6B Montane mesic meadows: 1 – rare occurrence
  • 6E Wet Cirsium meadows: 2 – optimum
  • 6F Intermittently wet Molinia meadows: 1 – rare occurrence
  • 7 Acidophilous grasslands

  • 7A Subalpine and montane acidophilous grasslands: 2 – optimum
  • 7B Submontane Nardus grasslands: 1 – rare occurrence
  • 11 Heathlands and scrub

  • 11H Subalpine deciduous scrub: 2 – optimum
  • 12 Forests

  • 12B Alluvial forests: 2 – optimum
  • 12F Limestone beech forests: 1 – rare occurrence
  • 12P Peatland pine forests: 1 – rare occurrence
  • 12Q Peatland birch forests: 1 – rare occurrence
  • 12R Acidophilous spruce forests: 1 – rare occurrence
  • 12S Basiphilous spruce forests: 1 – rare occurrence
  • Constant taxon

    ?
  • Constant taxon of associations: RBC04 Bartsio alpinae-Caricetum nigrae
  • Ecological specialization indices

    ?
  • Ecological specialization index for all vegetation types: 4.9
  • Ecological specialization index for non-forest vegetation: 5.4
  • Ecological specialization index for forest vegetation: 5.2

Distribution and frequency

  • Floristic zone: boreal, northern temperate, southern temperate, submeridional
    ?
  • Floristic region: Europe, Siberia
    ?
  • Elevational belt in the Czech Republic: colline belt, submontane belt, montane belt, subalpine belt
    ?
  • Occurrence frequency in the basic grid mapping cells and quadrants of the basic grid mapping cells: 380, 815
    ?
  • Commonness in vegetation plots from the Czech Republic

    ?
  • Occurrence frequency in vegetation plots: 0.3 %
  • Occurrence frequency in vegetation plots with a cover above 5%: 1.9 %
  • Occurrence frequency in vegetation plots with a cover above 25%: 0 %
  • Occurrence frequency in vegetation plots with a cover above 50%: 0 %
  • Mean percentage cover in vegetation plots: 2 %
  • Maximum percentage cover in vegetation plots: 13 %
  • Number of habitats with taxon occurrence in the Czech Republic

    ?
  • Number of narrow habitats in which the taxon occurs: 19
  • Number of narrow habitats in which the taxon has its optimum: 5
  • Number of broad habitats in which the taxon occurs: 7
  • Number of broad habitats in which the taxon has its optimum: 5
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